Reigechimys octodontiformis gen. et sp. novo and R. plesiodon sp. novo are described. They represent the frrst record of the family Echimyidae for the Cerro Azul Formation (Huayquerian Age, Late Upper Miocene) at La Pampa Province, central Argentina. Both species have hypsodont cheek teeth with an eight-shaped occlusal design. This dental morphology represents a noticeable case of convergence to octodontids and indicates that these echimyids inhabited open environments. 

Palaeotis weigelti, from the Middle Eocene of central Europe, is a flightless, paleognathous bird. It appears to be a member of the ostrich lineage on the basis of trivial derived characters. It is a very primitive ratite, however, and does not possess any of the highly specialized cursorial adaptations that characterize the modern steppe -and savanna- dwelling ostriches. 

Equus caballus antunesi, nova subspecies, was a hypsodont, slender, and rather small horse (around 141cm at the withers), with narrow hooves and protocones longer on P3/-P4/ than on M1/-M2/. It does not fit in any of the different "types" of Pleistocene caballine horses previously recognized but may be related to the horse from the Acheulean of Solana del Zamborino.
Hypsodonty, small size, slenderness, narrow hooves are all characters that can easily be related to ecological conditions. Equus caballus antunesi was probably a horse adapted to rather dry and cold conditions and to a hard ground. It does not seem related at all to the North-European Equus caballus germanicus-gallicus group. 

New and hitherto unpublished mammals from the stratigraphical levels Unterkohle, Untere Mittelkohle and Obere Mittelkohle of the Geiseltal near Halle, GDR, are described (= biochronological levels MP 11-13, Geiseltalian sensu Franzen & Haubold 1986a, b). The marsupial taxa Amphiperatherium aff. maximum (MP 12), A. goethei (MP 12), and Peratherium aff. monspeliense (MP 12 and 13) are recorded for the first time. A lectotype for Amphiperatherium giselense is designated, and the alleged primate Microtarsioides voigzi is assigned to Marsupialia, incertae sedis. A new insectivore species, Saturninia ceciliensis n. sp., is described (MP 13). The anteater Eurotamandua joresi is recorded for the first time outside its type locality, Grube Messel, FRG (MP 11). The present humerus and ulna display the autapomorphic features of the myrmecophagids. 

Editorial for celebrating the publication of the volume 20.

La revue PALEOVERTEBRATA a été fondée en 1967 par Louis THALER, à une époque où il n'existait, au plan international, que deux périodiques uniquement consacrés aux vertébrés fossiles. Actuellement ce nombre n'est que de quatre. A l'origine la revue fut essentiellement créée pour diffuser les résultats des chercheurs du Laboratoire de Paléontologie de l'Université de Montpellier, alors en début d'expansion et plusieurs thèses et de nombreux travaux directement liés à des thèses y furent publiés ce qui n'excluait déjà pas la publication d'articles d'auteurs étrangers.
[...] 

The study of dental variations in Landenodon from Dormaal (early Eocene, Belgium) shows that only one
species is present in this locality : L. woutersi QUINET, 1968 (Condylarthra, Arctocyonidae, Arctocyoninae). A high morphological variability is described, especially concerning the metaconid of P/4, the paraconid of M/2, and some characters of M1/ which have been found to be independant of each other. This variability is discussed. 

The geomyoid genus Dikkomys is well represented in a sample from the Black Bear Quarry Il local fauna of Early Hemingfordian age in Bennett County, South Dakota. Isolated unworn P/4's of Dikkomys matthewi WOOD have a prominent median cristid (sagicristid) with a connection to the metaconid and the hypolophid. With wear, P/4 does not become as molariform as P/4 because of this cristid.
A large sample of the Whitneyan beteromyid Proheteromys nebraskensis WOOD contains variants of the P/4 with on incipient sagicristid in approximately 18 percent of the population. The upper dentition and lower molars of Proheteromys nebraskensis are sufficiently generalized to indicate probable ancestry to Dikkomys. 

La découverte de nouveaux gisements à micromammifères dans le Miocène du Sud de la France et d'Espagne montre que, s'il n'est plus possible d'accepter dans son intégralité le schéma phylétique de Fahlbusch du genre Megacricetodon, il existe toutefois des segments de lignées à l'intérieur de provinces géographiques différentes (Péninsule Ibérique, France, Europe Centrale), avec lesquels on peut établir une biochronologie locale, segments qu'il est aussi possible de placer les uns par rapport aux autres, en utilisant les corrélations que l'on peut reconnaître entre certains niveaux repères des mammalogistes et l'échelle stratigraphique marine.
En utilisant ces données nouvelles, et d'après l'étude de la première molaire inférieure, nous proposons le
schéma évolutif suivant, pour les espèces de grande taille :

1 - au début du Burdigalien supérieur. une espèce commune, M. collongensis, issue de la forme ancestrale M. primitivus, est présente, de la Péninsule Ibérique à l'Europe Centrale. Son évolution est mal connue en Espagne, par contre cette lignée s'éteindrait en France au Burdigalien terminal avec le stade M. gersii nov. sp. de Sansan et de Luc-sur-Orbieu.
2 - au cours de la régression post-burdigalienne. reconnue en Languedoc et dans la Péninsule Ibérique apparaîtrait une espèce immigrante, M. crusafonti qui, dans cette dernière province, évolue pour donner M. ibericus de Nombrevilla. L'espèce M. gregarius de La Grive M et de La Grenatière dériverait de cette forme immigrants, en ayant une évolution différente.
3 - en Suisse, Bavière et Europe centrale, la lignée M. primitivus - M. collongensis va s'individualiser, dès le Burdigalien supérieur ou terminal, et donner naissance à la lignée M. bavaricus qui s'éteindra avec le stade M. germanicus nov. sp. du gisement anté - Hipparion de Anwil. Cette dernière espèce a été trouvée pour la première fois en France, dans le gisement de Collet Redon. 

     The Faunas of Rodents from seven north-african fossiliferous beds distributed from the Middle up to the Uppest Miocene are studied. One genus, seventeen species, one subspecies described are new.
       A detailed description is given of the evolution of the numerous specific lines, systematic groups and communities. The evidence of the strong endemism of the North African faunas of that time compared with those of Europe as well as of Tropical Africa is clearly shown. Special relationships with Asia clearly also appear, mostly in the Middle Miocene and the beginning of the Upper Miocene.
       Using the specific lines with a biostratigraphic purpose gives the opportunity to establish a biochronological scale for the continental formations of the Maghreb. This scale is calibrated by a few absolute datations. These datations, as well as the stratigraphical study of several fossiliferous localities have given the possibility to establish precise correlations with the corresponding scales of Europe and of Tropical Africa, as well as with the marine formations of the Neogene of occidental Mediterranean sea.
       The paleoclimatic data connected with this period in North Africa are explained and the contribution of the micromammals is discussed. 

La poche de Ste-Néboule, commune de Béduer (Lot), 15 km environ à l'WSW de Figeac, fait partie du groupe le plus septentrional des gouffres creusés par les ruissellements du Paléogène dans les calcaires jurassiques de la bordure sud-ouest du Massif Central et qui furent comblés à la même époque par des argiles sidérolithiques accompagnées de phosphate de chaux concrétionné ainsi que des restes de la célèbre faune dite «des phosphorites du Quercy» . 

The few remains recently collected at Ste Néboule (upper Eocene) belong to three genera: the Testudinid Ergilemys, of which it is the oldest occurence in Western Europa, probably Palaeochelys and an unpublished genus, which is peculiar to the Phosphorites du Quercy. Some statements are made on the chelonian fauna as a whole in that région. After the study of some characters proper to the genus Ergilemys, some remarks are made concerning its antecedents and descendants. 

The small insectivorous placental mammals from Ste-Néboule are poorly varied. Two orders of insectivores are documented : the Proteutheria by Pseudorhyncocyon cayluxi (leptictids), the Lipotyphla by Saturninia gracilis and S. beata (nyctitheriids). The bats are only rhinolophoids and mostly hipposiderids. Sufficient populations allow us to restrict the original definitions of three previous species (Hipposideros schlosseri, Palaeophyllophora quercyi, P. oltina). From now, these species can be regarded as monophyletíc. 

Le Quercy est aujourd'hui un vaste plateau calcaire, parcouru par un réseau karstique actif, pro· fondément entaillé par des vallées aux falaises abruptes, comme celles du Lot ou du Célé. Sur un sol peu épais domine la forêt de chênes, accompagnés de cornouillers, érables, genévriers. La faune est pauvre, peu diverse, et les nombreux chasseurs se satisfont de gibier d'élevage ...
Il y a trente-cinq millions d'années environ, le paysage était bien différent. La période de l'Eocène supérieur, qui s'achevait, avait été chaude et humide, si l'on se réfère à la fois aux paléotempératures (calculées à partir de sédiments marins extra-européens) et aux restes fossilisés de végétaux typiquement tropicaux.
Le Causse du Quercy devait être un plateau très disséqué par la karstification, à surface lapiazée creusée de gouffres en rapide évolution interne, et couvert d'un sol assez épais. Une forêt tropicale humide, avec notamment des Myricacées et des gymnospermes, recouvrait l'ensemble du pays, à peine interrompue au niveau des rares points d'eaux situés dans les bas-fonds. Dans ce cadre vivait une faune considérablement différente de la maigre faune actuelle. Si nous pouvons l'imaginer, c'est grâce aux cadavres des animaux entraînés dans les cavités par les eaux courantes ou les prédateurs, ou bien logeant et mourant dans les milieux souterrains. Nombre de ces restes, fossilisés, sont parvenus jusqu'à nous et sont aujourd'hui l'objet d'étude. 

The status of the giant bird taxa Liornis floweri and Callornis giganteus from the Santa Cruz Formation (late Early Miocene) of Patagonia, first described by Ameghino (1895) is reassessed on the basis of a re-examination of the type material at the Natural History Museum, London. Liornis floweri, which lacks a Pons supratendineus on the tibiotarsus and has an unbifurcated Canalis interosseus distalis on the tarsometatarsus, is clearly a brontornithid and is considered as a junior synonym of Brontornis burmeisteri. Ameghino’s replacement of Callornis by Eucallornis is unjustified. Callornis giganteus is a chimera based on a phorusrhacid tarsometatarsus (probably belonging to Phorusrhacos longissimus) and a brontornithid tibiotarsus. The latter can be considered as the lectotype of Callornis giganteus, which may represent a small morph of Brontornis burmeisteri or a distinct taxon. It is referred to here as Brontornithidae indet. The tarsometatarsus described by Dolgopol de Saez (1927a,b) as Liornis minor and considered by her as a gracile brontornithid apparently has a bifurcated Canalis interosseus distalis and should therefore be placed among the Phorusrhacidae. 

A large avian femur recently discovered at the Late Cretaceous Montplo-Nord locality at Cruzy (Hérault, southern France) is referred to the giant bird Gargantuavis philoinos. The estimated mass of the bird is 57 kg, within the range of living cassowaries. The specimen provides new evidence about the anatomy of G. philoinos, notably showing that the distal end of the femur was similar to that of modern birds in having a condylus lateralis subdivided into two semicondyles. A new diagnosis of Gargantuavis philoinos is provided and the taxon is placed in a new family of basal ornithurines.
  

The origin and early evolutionary history of the dinosaurs is a topic that has recently gone through a period of renewed interest and academic debate. For 130 years, one way of classifying the various dinosaur subgroups persisted as the accepted model, with increasing levels of research in the past quarter-century also providing evidence for the hypothesis that dinosaur origination occurred in the Southern Hemisphere, particularly in South America. It is, after all, from within the Late Triassic strata of countries like Argentina and Brazil that we get some of the very best early dinosaur specimens; many of these specimens are the earliest known representatives of some of the major dinosaur subgroups, such as the theropods and sauropodomorphs. However, some recent analyses have brought about a shift in terms of what is currently accepted and what is now disputed regarding the origin of dinosaurs – the Southern Hemisphere origination hypothesis was questioned (although this was based upon observations and not with quantitative analysis techniques), as has the shape of the dinosaur tree. Responses to the new hypothesis were numerous; many further supported a Southern Hemisphere point of origin. Whilst the interrelationships between the major dinosaur clades remains to be resolved, the current data does seem to comprehensively answer the question of where the dinosaurs first originated. However, it is arguable whether the current data that is being used in such palaeobiogeographical analyses is sufficient to provide an answer to the question of where specifically the dinosaur clade first appeared. This short communication urges a degree of caution about the current consensus and what steps may need to be taken to ensure that more meaningful results are produced in the future. 

Herein we describe a small but relatively diverse assemblage of fossil fishes derived from the lower Oligocene (Rupelian) Red Bluff Clay at site AMo-9 in Monroe County, Alabama, USA. Identified amongst the remains are 15 unequivocal taxa representing 11 families within five orders, and one additional taxon represents an unknown order and family. Taxa identified include Eostegostoma sp., Otodus (Carcharocles) sp., Mitsukurinidae/Carchariidae indet., Macrorhizodus praecursor, Galeorhinus sp., Negaprion gilmorei, Physogaleus sp., “Sphyrna” sp., Galeocerdo sp., cf. “Aetobatus” sp., Sphyraena sp., Xiphiorhynchus kimblalocki, Xiphiorhynchus sp., Cylindracanthus ornatus, and C. rectus. Several additional fossils could not be identified beyond Lamniformes, Carcharhiniformes, and Teleostei, but they likely belong to one of the identified taxa within this paleofauna. All of the fishes previously reported from the Red Bluff Clay within the entirety of the Gulf Coastal Plain of the USA are otolith-based, and each of the 15 unequivocal taxa reported herein are important new records for this lithostratigraphic unit. In particular, the Eostegostoma sp. and Xiphiorhynchus spp. specimens represent the first occurrences of these taxa in Alabama. The specimens of C. ornatus, Eostegostoma sp., and X. kimblalocki are stratigraphic and temporal range extensions from the middle and late Eocene into the Rupelian Stage of the Oligocene. Other described taxa may represent transitional forms between those described from the late Eocene and late Oligocene within the region. This study provides a tantalizing preliminary view into faunal transitions that occurred amongst marine fishes across the Eocene/Oligocene boundary within the Gulf Coastal Plain of the USA. 

Anthracotheres from the Pondaung Formation, Myanmar, are considered as one of the most primitive artiodactyl groups and they represent the oldest known record in the world. Thus, the understanding of this group has numerous implications for evolutionary biology and biochronological correlations. However, the systematlcs of these mammals has been interpreted in different ways, and the main debate focuses on the number of taxa represented in the Pondaung Formation. The revised taxonomy proposed here is mainly based on the relative development of the upper molar W-shaped ectoloph, system of crests and stylar cusps, and on body size. On the basis of these characters, they are classified into four genera including six different species. Two well-known genera, Anthracotherium and Anthracokeryx, are validated and more precisely diagnosed. Anthracokeryx possesses a better developed W-shaped ectoloph, system of crests and stylar cusps than Anthracotherium, which displays notable differences with the more derived representatives of this genus. Both of these Pondaung genera show evidence for sexual dimorphism. However, the incompleteness of fossil material fueled a debate concerning the status of two additional Pondaung anthracotheres, Siamotherium and Anthracohyus. The latter genus is of uncertain affinities, but it has been considered as a hippopotamid ancestor. Despite new material attributed to these two forms, additional discoveries are still required to establish their taxonomic status. The hypothesis that Southeast Asia was the centre of origin of Anthracotheriidae is supported by the retention of numerous primitive dental characters in these taxa and by the antiquity of the Pondaung Formation, to which an age of 37 My is now generally accepted. 

The knowledge of the Anchilophini has been lately renewed by the discovery of a rather large amount of new material still largely unpublished. This new material offers the opportunity of a systematic revision of this tribe gathering those of European Eocene Equoidea which bear no mesostyle on upper check teeth and display a heavy trend to the molarization of premolars.

A cladistic analysis has made out two genera, Anchilophus (Paranchilophus included as a subgenus), characterized by a marked lophodonty and the transverse narrowness of the cheek teeth, a rather high hypsodonty, the frequent occurrence of "crochets" and "anticrochets" on the superior ones, and a rather weak molarization of the premolars, opposite to Metanchilophus n. gen. whose cheek teeth are more transversally elongated, less high, less lophodont, with cusps better distinct, enamel thicker and premolars more molarized on the whole.

Three species of Anchilophus are recognized, A. desmaresti, type species of the genus, A. (Paranchilophus) remyi and A. (Paranchilophus) jeanteli n. sp.

The genus Metanchilophus is more diversified with the species dumasi, radegondensis, gaudini (whose a new sub-species M. g. fontensis is defined), depereli, castrensis n. sp. and chaubeti n. sp.

The skull anatomy has been moreover described with several taxa; it brings to light (for all that one can generalize) that Anchilophini were light animals with a slender and elongated snout, a thin zygomatic arch, a rather developed encephalon with an advanced gyrencephaly.

The structure of the nasal opening together with the occurrence of epitympanic sinuses and the molarizing process of the premolars corroborate the attribution of this tribe to the family PalaeotheIiidae. 

A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian.