The Late Triassic locality of Saint-Nicolas-de-Port (Meurthe-et-Moselle, France) has yielded numerous isolated teeth belonging to archosauriform reptiles. The following tooth groups can be identified: heterodont phytosaurs, the pterosaur Eudimorphodon, the prosauropod dinosaur Plateosaurus, three types of putative ornithischian teeth and 13 types of carnivorous Archosauriformes indet. Apparent venom-conducting teeth belonging to a new taxon of ?Archosauriformes (Graoullyodon hacheti nov. gen. nov. sp.) are also described. From a palaeogeographical point of view, the ornithischian teeth from Saint-Nicolas-de-Port (if their attribution is confirmed) are the oldest fossils of this group in Europe. The biostratigraphic distribution of the tooth forms mostly suggests a Late Norian or Early Rhaetian (depending on current interpretations) age of the deposits, but do not provide more precisions than fossils previously described from the area. The dietary habits and, consequently, the palaeoecological relationships of the different vertebrate groups discovered at Saint-Nicolas-de-Port are tentatively established: the ornithischian and prosauropod teeth reflect a herbivorous diet, whereas the other archosauriform teeth are probably from camivores or omnivores.

  

Carolocoutoia ferigoloi gen. et sp. nov. is the largest of protodidelphid marsupials, known from Middle Paleocene levels at Itaboraí Formation, southeastern Brazil. It differs from other members of this family in having molars with low cusps which are basally inflated, rounded crests without cutting edges, and a thick enamel layer which is wrinkled, specially at the labial half. A comparative analysis among representatives of this family led us to recognize only three genera undoubtely assignable to it:  Protodidelphis PAULA COUTO, 1952, Robertbutleria MARSHALL, 1987, and Carolocoutoia gen. nov. Protodidelphids lack the basic derived features diagnostic of Polydolopimorphian marsupials, while most of its derived features agree with its belonging to the Didelphimorphia. Protodidelphids comprise a specialized clade of opossum-like marsupials adapted to frugivorous or frugivore-omnivorous feeding habits. They differ from other didelphimorphians in having very large, spire-like entoconids, reduced and antero-posteriorly compressed paraconids, absence of stylar cusp C and of para- and metaconules, large stylar cusps B and D which are proximate to each other, short postmetacristae, eccentric protocones, and molars that increase rapidly in size from M/ml to M/m3.
  

THE ANIMALS. Systematische Zoologie by A. REMANE, V. STORCH, and U. WELSCH. 1986, Edition 3. Cloth and paperbound. Stuttgart and New York : Gustav Fischer Verlag. xvi + 698 pp. DM 86. (Geb.) & DM 72 (kart.). 

The description of new material of the Upper Eoœne of France, Southem Germany and Spain together with the restudy of the material described by Thaler (1966b) as Pairomys allows to confirm the validity of the genus Ectropomys up to know controversally discussed. As a consequence the subfamily Remyinae and Oltinomyinae can be defined precisely. The study of the dental characters shows that the Remyinae in fact belong to the Theridomyidae and represent a derived evolutionary unit within this family. The Oltinomyinae together with the Theridomyinae and Issiodoromyinae very probably form a monophyletic unit. 

New Dendromus species (Rodentia, Muroídea) from Langebaanweg (Pliocene, South Africa). Stratigraphical and paleoecological consequences.

Two new species of Dendromus are described from the Langebaanweg site which precises the evolutionary trend among this genus in South Africa and gives further paleoenvironmental indications. Two evolutionary stages are described: D. darti nov. sp. shows low-crowned molars with bunodont cusps and its more closest relative would be D. melanozis from the Cape region. On the contrary, D. averyi nov. sp. is more lophodont and is better related with the modem D. melanotis. Both species are at a less evolved stage than the Dendromus sp. from Laetolil Beds at Laetoli. The Langebaanweg deposits cannot still be dated by biostratigraphy but they clearly cannot be older than the basis of Pliocene times. The association of Dendromus and Mystromys in the same levels indicates a grassland environment with woodland patches as well as probable swamps. 

The St-Dézéry local fauna (3 reptile-, 4 mammal species) is approximately of the same age as the La Débruge or the Ste-Néboule faunas. It conduces to a better dating of the limestones underlying the Célas sandstones. A large part of a mandible of Amphimeryx was found there, which documents the record of this family of small artiodactyls 

The Laprade fauna is chronologically situated between those from Egerkingen and Lissieu and consequently, is close to the MP 14 reference-level of the European mammalian biochronological scale (Symposium of Mainz, 1987).
This new fauna is presently the oldest known in the Quercy phosphorites, formerly the Le Bretou fauna (MP 16) was considered as the oldest one. The Laprade fauna includes 21 species which belong in 7 mammalian orders (Marsupialia: Amphiperatherium bastbergense, Amphiperatherium goethei; Apatotheria: Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp.; Lipotyphla: Saturninia cf. mamertensis, Saturninia cf. intermedia; Chiroptera: Vespertiliavus lapradei nov. sp.; Rodentia: Protadelomys cf. lugdunensis, Elfomys nov. sp.; Primates: Nannopithex cf. filholi, cf. Pseudoloris or Pivetonia; Perissodactyla: ?Palaeotherium ?castrense, small-sized Palaeotherium sp., Plagiolophus sp., Anchilophus sp.; Artiodactyla: Dichobune cf. robertiana, Mouillacitherium cartieri, Tapirulus cf. depereti, Mixtotherium priscum, Pseudamphimeryx schlosseri, and Artiodactyla indet). Sixteen of these species are mentioned for the first time from the Quercy faunas.
The recognition of a new apatemyid, Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp., attests to an early origin of a lineage known in the Late Eocene. The emballonurid bat Vespertiliavus lapradei nov. sp. is presently the earliest record of this genus and family. This Auversian fauna leads to discuss the age of taxa showing archaic features. These taxa were defined on specimens collected in the Quercy during the last century and have never been found in the Quercy localities recently investigated. This fauna bears also evidence of a karstic filling episode older than those previously dated by fossils in the Quercy Jurassic. 

Editorial for celebrating the publication of the volume 20.

La revue PALEOVERTEBRATA a été fondée en 1967 par Louis THALER, à une époque où il n'existait, au plan international, que deux périodiques uniquement consacrés aux vertébrés fossiles. Actuellement ce nombre n'est que de quatre. A l'origine la revue fut essentiellement créée pour diffuser les résultats des chercheurs du Laboratoire de Paléontologie de l'Université de Montpellier, alors en début d'expansion et plusieurs thèses et de nombreux travaux directement liés à des thèses y furent publiés ce qui n'excluait déjà pas la publication d'articles d'auteurs étrangers.
[...] 

Is hereby described an impressive lchnoiauna belonging to the Lower to Middle Ludian of the Gard (S. France). The slab, already cleaned over a length of 18 m, is located near the top of the Potamides aporoschema lacustrine limestone (Lower Ludian, Euzet zone). It is therefore older than the Célas sandstone deposit, and still more than the Melanoides albigensis and M. acutus marly limestone corresponding to the Upper Levels of the Ludian stage. Although biostratigraphically older than the La Débruge and Montmartre zone, the biotope shows already a sampling of very tall Artiodactyles, Perissodactyles and Carnivorous. One of the most « majestic ›› Artiodactyles, Anopolotheriipus lavocati, nov., points out a huge size type. To mention also among the Ichnotypes described, 10, the big Perissodactyle Palaeotheriipus similimedius, nov., and the big Carnivorous Hyaenodontipus praedator, nov. 

New dental and skeletal material referable to Palaeochiropteryx tupaiodon from the Middle Eocene locality of
Messel (G.F.R.) is studied, which provides additions to the previously gained knowledge of this european genus. Dental specimens from Geiseltal (G.D.R.), also of Middle Eocene age, allow us to analyze Cecilionycteria prisca. Some of these are separated to establish a new genus, Matthesia, and two new species, M. germanica and M. ? insolita. 

The study of dental variations in Landenodon from Dormaal (early Eocene, Belgium) shows that only one
species is present in this locality : L. woutersi QUINET, 1968 (Condylarthra, Arctocyonidae, Arctocyoninae). A high morphological variability is described, especially concerning the metaconid of P/4, the paraconid of M/2, and some characters of M1/ which have been found to be independant of each other. This variability is discussed. 

The wear facets of primitive rodents can be homologized with those of primitive primates and ungulates. As in primates, the jaw movement was ectental, with an increased anterior component in the lingual phase (phase ll). The buccal phase (phase I) in rodents approaches the horizontal and it tends to be reduced in importance in comparison with the lingual phase. ln more advanced rodents the efficiency of grinding is increased by the development of additional cutting edges of enamel (e.g. enlargement of hypocone, development of mesoloph and lingual sinus). The buccal phase movement becomes lined up with the lingual phase movement to form a single oblique chewing stroke,resulting in planation of the crown. As the stroke becomes more longitudinal (propalinal) the enamel edges become more transverse. In Muridae propalinal chewing evolved before the loss of cusps, facets were reorientated and additional cusps developed. 

The Arvicolid evolution results in an increase of the dental structure complexity. The M3/ differenciation seems to characterise the tribe subdivisions, that of M/1 being variable from one to another lineage. The phyletic relationships of fossil lineages are discussed from a paleogeographic point of view. 

A well-preserved anterior cranium of Paranourosorex gigas RZEBIK-KOWALSKA, 1975 from Podlesice, Poland, provides additional knowledge of this rare shrew, including almost complete maxillary dentition. The systematic status of this species as a relative of recent Anourosorex MILNE-
EDWARDS, 1870 is confirmed 

The Caudata are known by two Salamandridae ; one of them is attributed to the genus Megalotriton. The Pelobatidae form the major part of the Anura ; a few bones indicate also the presence of Neobatrachia. 

The few remains recently collected at Ste Néboule (upper Eocene) belong to three genera: the Testudinid Ergilemys, of which it is the oldest occurence in Western Europa, probably Palaeochelys and an unpublished genus, which is peculiar to the Phosphorites du Quercy. Some statements are made on the chelonian fauna as a whole in that région. After the study of some characters proper to the genus Ergilemys, some remarks are made concerning its antecedents and descendants. 

Crocodilians are represented in the Upper Eocene of Sainte·Néboule (Lot) by an isolated parietal and a dorsal scute, bath from young individuals. They are refferred to Diplocynodon sp. Predators (mammals and birds) are probably responsible for the occurrence of remains of small crocodilians (belonging to the genera Allognathosuchus and Diplocynodon) in the phosphorltes of Quercy. 

La poche à phosphorite de Sainte-Néboule (Lot) a livré au cours des récentes campagnes de fouilles effectuées dans le cadre de la RCP 311 une douzaine de dents ou fragments de dents de paléothéridés à rapporter à 3 taxons. 

The Late Miocene site of Gritsev (MN 9, Ukraine) has yielded a very rich bat fauna, the remains of which are well preserved. Compared to other Neogene bat assemblages of Europe, the Gritsev bat community is unique in preserving plecotine bats, which are rare from Neogene sites. Some peculiar and new bat species, including a large plecotin Otonycteris, already were described from the Gritsev mammal site. Here we report new records of small plecotin bats from Gritsev, including a new taxon, Barbastella maxima nov. sp. This is the earliest reliable fossil record of this genus and it differs from more recent species of Barbastella in being considerably larger. The evolutionary patterns in the odontology within the tribe Plecotini, supported by biostratigraphical distribution of fossil records of Plecotus are discussed. The morphological peculiarities of the new fossils of plecotine bats from Gritsev are discussed in connection with its possible taxonomical affinity. 

The origin and early evolutionary history of the dinosaurs is a topic that has recently gone through a period of renewed interest and academic debate. For 130 years, one way of classifying the various dinosaur subgroups persisted as the accepted model, with increasing levels of research in the past quarter-century also providing evidence for the hypothesis that dinosaur origination occurred in the Southern Hemisphere, particularly in South America. It is, after all, from within the Late Triassic strata of countries like Argentina and Brazil that we get some of the very best early dinosaur specimens; many of these specimens are the earliest known representatives of some of the major dinosaur subgroups, such as the theropods and sauropodomorphs. However, some recent analyses have brought about a shift in terms of what is currently accepted and what is now disputed regarding the origin of dinosaurs – the Southern Hemisphere origination hypothesis was questioned (although this was based upon observations and not with quantitative analysis techniques), as has the shape of the dinosaur tree. Responses to the new hypothesis were numerous; many further supported a Southern Hemisphere point of origin. Whilst the interrelationships between the major dinosaur clades remains to be resolved, the current data does seem to comprehensively answer the question of where the dinosaurs first originated. However, it is arguable whether the current data that is being used in such palaeobiogeographical analyses is sufficient to provide an answer to the question of where specifically the dinosaur clade first appeared. This short communication urges a degree of caution about the current consensus and what steps may need to be taken to ensure that more meaningful results are produced in the future.