Prolagus sardus is the last representative of the diverse lineages of European endemic ochotonids. It is also the most abundant in the collections. The previous studies made of this species have established rather well its dental morphology, its phylogenetic position, its geographic and temporal distribution, and its intraspecific individual variation. On the other hand, no osteologic study has fully utilized the superb material from Corsica and Sardinia  collected by Forsyth Major.
Nearly all of the parts of the skeleton are represented in this material; they are here described and figured. Comparisons are made with Ochotona, the only surviving genus of the family, as well as with the living leporids, in particular Romerolagus, Oryctolagus and Lepus. A brief examination of the dentition reveals some particular characters of the incisors. The osteologic study allows limited interpretations to be advanced concerning the posture, mastication. locomotion and some other adaptive features of P. sardus.
P. sardus appears as an incontestable ochotonid, but it differs from Ochotona by some characters which are found in leporids. Certain of the characters are here judged primitive for lagomorphs. The genus Ochotona therefore can no longer be considered as the image of a primitive lagomorph, because in spite of the lack of cursorial adaptations which differentiate it from leporids, it presents other specializations which are common to it alone. 

A new example of parallelism in the dental pattern ofthe Theridomyidae is illustrated by the description ofa new species: Theridomys Iudensis from the standard level of Antoingt (middle Oligocene). Considering the occurence ofthis parallelism phenomenon. the use of numerous qualitative and quantitative criteria is essential to characterize the different stages ofthe different evolutive lineages. Thus, a new simple parameter is proposed (CHY = H+l/0,5 L) to estimate hypsodonty of the medium hypsodont Rodentia. 

Micro-CT imaging is an increasingly popular method in paleontology giving access to internal structures with a high resolution and without destroying precious specimens. However, its potential for the study of hypoplasia defects has only recently been investigated. Here, we propose a preliminary study to test whether hypoplastic defects can be detected with micro-CT (μCT) scan and we assess the costs and benefits of using this method instead of naked eye. To do so, we studied 13 fossil rhinocerotid teeth bearing hypoplasia from Béon 1 (late early Miocene, Southwestern France) as positive control and 11 teeth of the amynodontid Cadurcotherium (Oligocene, Phosphorites du Quercy, Southwestern France), for which enamel was partly or totally obscured by cement. We showed that all macroscopically-spotted defects were retrieved on 3D reconstructions and selected virtual slices. We also detected additional defects using μCT scan compared to naked eye identification. The number of defects detected using μCT was greater in the Cadurcotherium dataset (paired-sample Wilcoxon test, p-value = 0.02724) but not for our control sample (paired-sample Wilcoxon test, p-value = 0.1171). Moreover, it allowed for measuring width and depth of the defects on virtual slices (sometimes linked to stress duration and severity, respectively), which we could not do macroscopically. As μCT imaging is both expensive and time consuming while not drastically improving the results, we recommend a moderate and thoughtful use of this method for hypoplasia investigations, restricted for instance to teeth for which enamel surface is obscured (presence of cement, uncomplete preparation, or unerupted germs). 

The early gnathostome genus Mesacanthus is well represented in both Lower Old Red Sandstone and Middle Old Red Sandstone assemblages of northern and central Scotland. This ‘acanthodian’ taxon is currently thought to comprise two valid species: M. mitchelli and M. pusillus. Although the whereabouts of the holotype of M. mitchelli (NHMUK PV P560) is known, the syntype material for M. pusillus has long been thought lost. Here we identify at least one specimen that formed part of the original syntype material for M. pusillus, albeit in a slightly different condition than when it was originally figured. This specimen is ROM 25872, which is here designated as the lectotype. A second specimen – ELGNM 1978.191.1 – could represent another of the syntype specimens, but poor preservation quality makes it impossible to be certain. 

As a result of the enrichment of eotheriodont material by one of us (P.K.T.), these specimens (essentially Biarmosuchur and Eotitanosuchur) are reexamined and refigured. A reevaluation of their particularities supports the distinction of two families, for which new diagnoses are proposed. This leads us to discuss the affinities of these families, with respect to the sphenacodonts on one hand, and to the South African primitive theriodonts on the other (gorgonopsids and ictidorhinids). This study contains inherent paleogeographic consequences which are considered in conclusion. 

Les Pistes de Vertébrés du Stormberg Supérieur ("Trias terminal à Rhétien"), ou Quthingien

Si les zones du Stormberg inférieur se sont révélées contenir de nombreuses traces, surtout dans les faciès dits "Molteno moyen et supérieur", représentant apparemment la base du Keuper, il est frappant de voir pratiquement l'ensemble de cette grosse faune "Molteno" disparaître avec la fin de cette période, que nous avons appelée le "Maphutsengien".

Dès les premières zones du Stormberg supérieur, que nous nommons le"Quthingien" la zoocénose et la phytocénose, en même temps que les données d'ensemble manifestées par l'environnement, sont modifiées. Nous ne verrons plus guère de dépôts marécageux à flore riche et variée, parfois même luxuriante. Les fougères elles-mêmes ont disparu. Elles sont remplacées par de maigres plantes, aux feuilles très souvent filiformes qui paraissent témoigner d'un climat continental. Le sol est devenu de plus en plus rouge, avec des variations latérales beaucoup plus accusées. Les fleuves amenant des galets des monts du "Grand Sud" ont tari. La faune va en subir les conséquences. Certaines des espèces se révèleront sautillantes ou coureuses, pour un grand nombre plus légères et pour la quasi-totalité d'apparence carnivore ou entomophages, les phytophages devant se contenter d'un régime ingrat,difficile ou à tout le moins irrégulier,les dépôts le montrent.

C'est dans ces conditions que s'inaugure notre Etage nouveau,quelque peu discordant sur les zones A/5, A/6 ou A/7 du Stormberg inférieur (Maphutsengien). Le Stormberg supérieur (ou Quthingien) commence avec le paléopaysage remarquable dit de Moyeni, que nous allons maintenant étudier, typologiquement, avec ses homologues du même âge. Quelques 38 types d'animaux tous nouveaux vont défiler à nos yeux lors de la zone de base de cet Etage, ou zone B/1.

L'on nous avait proposé d'intituler ce Ile Tome de la série : "La grande Dalle de Moyeni et ses homologues. Paléo-spectacles, scènes et paysages animaux au Lesotho à l'approche du Trias finissant". Nous avons préféré garder le sous-titre plus haut, peut-être plus prosaïque.

Un llle Tome est en préparation : "Les développements ultérieurs et terminaux de la faune du Gondwana".

  

The earliest cetaceans are found in the early Eocene of Indo-Pakistan. By the late middle to late Eocene, the group colonized most oceans of the planet. This late Eocene worldwide distribution clearly indicates that their dispersal took place during the middle Eocene (Lutetian). We report here the first discovery of a protocetid fossil from middle Eocene deposits of Senegal (West Africa). The Lutetian cetacean specimen from Senegal is a partial left innominate. Its overall form and proportions, particularly the well-formed lunate surface with a deep and narrow acetabular notch, and the complete absence of pachyostosis and osteosclerosis, mark it as a probable middle Eocene protocetid cetacean. Its size corresponds to the newly described Togocetus traversei from the Lutetian deposits of Togo. However, no innominate is known for the Togolese protocetid, which precludes any direct comparison between the two West African sites. The Senegalese innominate documents a new early occurrence of this marine group in West Africa and supports an early dispersal of these aquatic mammals by the middle Eocene.
  

The Tafna Basin corresponds to the lowlands, which are located in front of Tessala and Traras ranges, below the Tlemcen mountains, Algeria. This basin displays a complete sedimentary cycle dominated by lagoonal-fluvial and marine deposits. The continental formations located at the base of these deposits are mainly composed of alternating sandstones and clays. An early late Miocene age has been previously attributed to them, based on direct correlations with marine deposits. Search for micromammal fossils led to the discovery of three different rodent species from a single level of the Djebel Guetaf section, located at the bottom of these deposits. The rodent assemblage indicates a late Miocene age. Combined magnetostratigraphical and biostratigraphical investigations were carried out to provide a more accurate age control of these continental deposits. Sixty-four oriented rock samples were collected for a magnetostratigraphic study along a 92 meters thick section including the fossiliferous layer. Rock magnetic investigations indicate the presence of both high and low coercivity minerals. Specimens subjected to progressive thermal demagnetization procedures show that the samples exhibit a high temperature magnetization component and display a normal polarity. Based on biostratigraphic constraints, the Guetaf section is correlated with Chron C4An, indicating an age ranging from
9.1 to 8.7 Myr.
  

Morphological and biometrical description of skulls, teeth, and limb bones of extant and fossil Old World herniones (including E. hydruntinus) and of New World 'stilt-Iegged' and other slender species from Blancan to Holocene. An Appendix presents ways in which the approximate size of some missing bones or dimensions may be deduced from available ones.

The discussed and/or illustrated fossils were found in Bolivia (Tarija), Canada (Yukon), China (Choukoutien, Gulongshan, Jiling, Loufangzi), Ecuador (Oil Fields), Ethiopia (Melka Kunturé), France (Lunel-Viel), Germany (Süssenborn), Greece (Agios Georgios, Petralona), Hungary (Dorog), Italy (Romanelli), Mexico (Cedazo, San Josecito), Mongolia (Sjara-osso-gol), Spain (Venta Micena), ex-Soviet Union (Akhalkalaki, Binagady, Chokurcha, Chukochya, Kabazi, Kolyma, Krestovka, Kurtak, Staroselie, Tologoj), USA (Alaska, Arkalon, Cedar Meadow, Channing, Conkling, Dry Mountains, Hay Springs, Leisey Shell Pit A, Lissie Formation, Natural Trap, Pool Branch, Powers Ranch, Rock Creek, San Diego, Santo Domingo, Seymour Formation, Shelter, Slaton, Trinity River). Numerous raw or statistically elaborated data are given in Tables.

There is no evidence for the existence of Old World hemiones in the New World nor of 'stilt-Iegged' equids in the Old World. The first 'stilt-Iegged' equid was found at Santo Domingo, New Mexico, and is believed to be Late Blancan. It was probably at the origin of E. calobatus (Arkalon, Rock Creek) and of the smaller E. semiplicatus (Channing, Rock Creek). Slender, but not 'stilt-Iegged', equids found at Natural Trap, Wyoming, ca. 12 ky ago belong to Amerhippus. AlI these species share with Oid World Sussemiones (and some hemiones) peculiar patterns on the lower cheek teeth.

The slender Equus sp. B of Leisey Pit A, Florida, ca. 1.2 Ma, as weIl as Amerhippus francisci and E. tau (probably a senior synonym of E. quinni) share conventional lower cheek teeth patterns. The skulls of A. francisci and E. tau, however, are quite different.

Paleontological data suggest a common origin of Amerhippus, Sussemiones, and 'stilt-Iegged' equids during the late Blancan. Old World hemiones seem to have differentiated later. 

The analysis of oxygene isotope variations as well as paleobotanical data suggest that the Oligocene/Miocene boundary corresponds to a transitional period marked by floristical and climatic variations. During this period, the pyreneo-alpine tectonics has contribued to modify the geography and western Europe landscapes. Faunal changes (appearances, extinctions, migrations) are observed in different mammalian groups, notably in the rodents. A study of the evolutionary trends and patterns in paleogene rodents is involved for the period ranging from level MP 28 of the Late Oligocene to the Early Miocene, including the Oligo-Miocene boundary.
The Rodents fauna from the sites of Venelles (Bouches-du-Rhône District, France) and Thezels (Lot, France), previously mentionned in litterature, have been studied. The first description of the Eomyidae of La Milloque (MP 29) has been completed. These faunas are compared to those from various localities dating from the considered period. In La Milloque, a new representative of the Eomys species is described next to a form close to Rhodanomys hugueneyae ENGESSER, 1987. It is the Eomys milloquensis nov. sp., the likely descendant of Eomys quercyi COMTE & VIANEY-LIAUD, 1987. Two new species are also described in Thezels: Eucricetodon thezelensis nov. sp., resulting from a likely and local evolution of Eucricetodon praecursor (SCHAUB, 1925) from La Milloque, which, in the same geographic area, could be at the origin of Eucricetodon hesperius ENGESSER, 1985 from Paulhiac. Plesiosminthus admyarion nov. sp., quite distinct from Plesiosminthus schaubi VIRET, 1926, which announces Plesiosminthus myarion SCHAUB 1930. Venelles 'Plesiosminthus schaubi population is considered as a sub-species, named Plesiosminthus schaubi meridionalis nov. subsp. New phylogenetic patterns are proposed. Among the Eomyidae, a quantification of various features of the M1-2/ crown (hypsodonty, degree of abrasion, occlusal angle, state of development of the I and V anticlines), and a comparison with the occlusal diagram of the other teeth among various other populations allows a more efficient separation of Eomys and Rhodanomys genera. In Western Europe, and within this period, it finally does not seem possible to gradually connect the genus Eomys to the genus Rhodanomys. The evolution of the Eomys quercyi - milloquensis lineage seems to underline a similar evolution to that which may have led from the Eomys to the Rhodanomys form. The latter which appears totally accomplished at level MP 29 of the Oligocene is considered as an immigrant. If we compare the most representative species of the Venelles, Thezels, and Coderet sites, (i.e. Rhodanomys, Eucricetodon, Adelomyarion, Peridyromys, Plesíosminthus), it becomes impossible to confirm their biochronological separation. The noticeable differences between the populations may be interpreted as geographical variations. An explanation to these variations, and to fauna's evolution during the Late Oligocene and Early Miocene can be found in the environmental modifications, supported by isotopic, paleobotanical and sedimentologic analysis. A tentative reconstruction of the environments is attempted by the cenogram method. The analysis of the fluctuations of fauna's diversity shows variations which may be correlated to a drop in temperature at MP 29, during the Late Oligocene, followed by an increase in temperature along with an aridity phenomenom, during the basal Miocene (MN O).The confrontation of various methods give the opportunity of reconstituting and comparing the evolution of the environment of three sequences of sites chosen from different regions. Ecological affinities of various rodents' species are being examined. It is possible to consider that the integration of all the conclusions resulting from this study should lead to an explanation to the evolution of rodents for the period around the Oligocene-Miocene boundary. The site of Coderet- level 3- would be posterior to the latter, at the beginnig of the Miocene, and would mark the level MN 0 of the Aquitanian.

  

This paper is an English concentrate of various Russian publications by the senior author presenting the mammaIian taxa from the Cretaceous (Albian through Santonian) of the region termed Middle Asia by Soviet geographers. The diagnoses are the unmodified, literal translation of the original version, but are followed with short complementary remarks; most of the species are illuslrated anew with SEM photographs, others are by normal photography. The fossiliferous formations are cited and arguments for their dating are given. Finally, the main vertebrate groups accompanying mammaIs are listed and the environment and climate at the time of deposition are suggested. In conclusion, an hypothesis on the origin and high diversity of tribosphenic mammals on the Cretaceous coastal plains of southwest Asia is proposed. In appendix the taxon Khuduklestes bohlini novo gen. novo sp. is formally defined. 

This paper presents a preliminary list of insectivores from the Vallesian beds at Montredon (France). The associated rodent fauna has established a Vallesian age for the fauna. Eleven species belonging to the Soricidae, Talpidae, Erinaceidae, and Dimylidae are identified of which four only are referred with certainty to forms already named. 

Eosimiid and amphipithecid primates document a long and significant history of primate evolution throughout the Eocene in Southeast Asia. Despite the absence of a comprehensive post-Eocene fossil record, it was generally hypothesized that both families left no descendant in Asia. Recently, two new small-bodied taxa, Bugtipithecus and Phileosimias, have been recovered in early Oligocene coastal deposits from the Bugti Hills (Balochistan, central Pakistan) and referred to the families Amphipithecidae and Eosimiidae, respectively, on the basis of dental fossil remains. In this paper, we provide more exhaustive description, comparison, and discussion of these taxa. As for tarsiid and sivaladapid primates, the persistence of eosimiids and amphipithecids into the Oligocene clearly demonstrates that low latitudes of South Asia provided a continuous access to tropical refugia during the climatic deterioration characterizing the late Eocene-early Oligocene interval, which was seemingly lethal for primate communities elsewhere across the Holarctic continents. As a contribution to the ongoing phylogenetic debates regarding the position of eosimiids and amphipithecids on the primate family tree, we have performed a cladistic analysis in a high-level primate systematic context in order to assess the position and the role of these new taxa in that phylogenetic issue. Our results support the view according to which eosimiids and amphipithecids (and by extension Phileosimias and Bugtipithecus, respectively) are stem anthropoids. These fossils from Pakistan document an unsuspected Oligocene phase of the evolutionary history of anthropoid primates in southern Asia, which clearly enhances the extent of the anthropoid radiation in this province during the Paleogene. Several phylogenetic and paleobiogeographic aspects are discussed, notably the intra- and inter-relationships between Paleogene Asian and Afro-Arabian anthropoids, and the resulting potential dispersal models between both land-masses during the Paleogene. 

This paper is mainly devoted to the systematics of rodents from two middle Eocene (Bartonian) localities: Chéry-Chartreuve and Rocourt-Saint-Martin (Aisne, Eastern Paris Basin). These two localities are stratigraphically located slightly above the Auversian sands. The two faunas, which comprise 11 and 8 taxa, respectively, are very different in their composition. That of Rocourt-Saint-Martin shows strong similarities with that of the geographically very close locality of Grisolles, referred to the MP16 mammalian Reference level. The very distinct fauna of Chéry-Chartreuve includes a new species of Ailuravinae, Ailuravus nov.sp, and some teeth of the theridomyid Protadelomys, which represent archaic elements in the fauna. The most abundant species of the locality represents a new genus of primitive Theridomyidae. The presence of some teeth belonging to a new species of large Remyinae, Remys nov. sp., of Elfomys engesseri HOOKER & WEIDMANN, and a population of small dimensions referred to the genus Estellomys allow a correlation with Les Alleveys (Switzerland), with however some differences that would indicate an older age for Chéry-Chartreuve. Situated at the base of the "Marinesian" from the Bassin de Paris, this fauna is unquestionably different from those referred to the MP16 reference level and could represent an older level for which the macrofauna remains very poorly known. Conversely, the comparison of rodents from La Livinière II with those present in MP16 faunas, especially those of Robiac (Gard), shows a great similarity between both localities. This casts doubts on whether to keep this La Livinière II faunule to define the current MP15 reference level, while the biostratigraphical position of Pontils 26 (Spain), previously referred to this level, is reconsidered. Chery Chartreuse could be a good candidate for a new definition of the MP15 reference level. 

The present work is devoted to the study of the Cretaceous and Tertiary fishes (teeth of Selachians, Actinopterygians and Dipnoans) collected during a recent expedition in Niger. The Maestrichtian localities have yielded a new genus and a new subspecies of Selachian: Igdabatis sigmodon nov. gen., nov. sp. and Lamna biauriculata nigeriana nov. subsp. The locality of Sessao, which has been attributed to the Thanetian by means of the study of the fish, has furnished by screen-washing an interesting fauna wherein six new species are described: Raja Iouisi, Dasyatis sessaoensis, D. sudrei, D. russelli, Hypolophites thaleri and Ceratodus casieri. Comparison of these faunas with contemporary faunas of Africa has brought out a certain endemism in the Iullemmeden Basin during the late Cretaceous and the early Tertiary. 

The knowledge of the Anchilophini has been lately renewed by the discovery of a rather large amount of new material still largely unpublished. This new material offers the opportunity of a systematic revision of this tribe gathering those of European Eocene Equoidea which bear no mesostyle on upper check teeth and display a heavy trend to the molarization of premolars.

A cladistic analysis has made out two genera, Anchilophus (Paranchilophus included as a subgenus), characterized by a marked lophodonty and the transverse narrowness of the cheek teeth, a rather high hypsodonty, the frequent occurrence of "crochets" and "anticrochets" on the superior ones, and a rather weak molarization of the premolars, opposite to Metanchilophus n. gen. whose cheek teeth are more transversally elongated, less high, less lophodont, with cusps better distinct, enamel thicker and premolars more molarized on the whole.

Three species of Anchilophus are recognized, A. desmaresti, type species of the genus, A. (Paranchilophus) remyi and A. (Paranchilophus) jeanteli n. sp.

The genus Metanchilophus is more diversified with the species dumasi, radegondensis, gaudini (whose a new sub-species M. g. fontensis is defined), depereli, castrensis n. sp. and chaubeti n. sp.

The skull anatomy has been moreover described with several taxa; it brings to light (for all that one can generalize) that Anchilophini were light animals with a slender and elongated snout, a thin zygomatic arch, a rather developed encephalon with an advanced gyrencephaly.

The structure of the nasal opening together with the occurrence of epitympanic sinuses and the molarizing process of the premolars corroborate the attribution of this tribe to the family PalaeotheIiidae. 

Abstract: A very tiny cuckoo, Chambicuculus pusillus, was recently described after a few fragments of tarsometatarsi from the late Early/early Middle Eocene of Djebel Chambi, Tunisia. New remains, notably a coracoid, confirm the attribution of this genus to the recent family Cuculidae. This coracoid shows a very elongate and strap-like processus procoracoideus. This morphological feature is otherwise only known in the family Cuculidae. The characteristics of the coracoid and tarsometatarsi show that Chambicuculus is morphologically more advanced over the other stem cuculids described in Europe and North America. Chambicuculus is the oldest Cuculidae known so far. 

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

The biometric and morphologie variability of the cheek teeth in the end-of-the-phylum species Lophiodon lautricense Noulet studied in this note, reposes on the observation of about 800 teeth. These were revealed to be little variable in absolute dimensions. The considerable morphologie variability in the upper premolars permitted the problem of the molarization process to be taken up. An hypothesis concerning the order of eruption of the cheek teeth is formulated based on an examination of a large number of milk dentitions. In conclusion, it is suggested that reservations be held on the value of dental characters classically used in systematics for the group under consideration. 

Bulk-sampling of fossil-rich tempestites from the Chattian Sülstorf Beds of
Mecklenburg, north-eastern Germany, yielded a rich selachian fauna in which batoids
predominate by the abundance of teeth but are subordinate by the number of taxa. Thirteen
taxa are identified, among which rajiform batoids are the most diverse (six species). One
genus and three species are newly described: Raja thiedei sp. nov., Oligoraja pristina gen. et
sp. nov., and Torpedo chattica sp. nov. Two species are reallocated: Atlantoraja cecilae
(Steurbaut & Herman, 1978) new comb., and Dipturus casieri (Steurbaut & Herman, 1978)
new comb. Ontogenetic heterodonty is documented for the first time in the dental pattern of
Myliobatis sp. Stratigraphical ranges of batoid taxa in the period from Rupelian to Langhian
are presented and partly discussed in context with the palaeoclimatic evolution and
palaeogeographic situation of the North Sea Basin.