Abstract not available 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

Confident attribution of bipedal tridactyl dinosaur tracks to theropods or ornithopods can be challenging. Here we describe trackways produced by tetanuran dinosaurs, previously attributed to hadrosaurs, from Coahuila State, northeastern Mexico. Multiple trackways headed in the same direction suggest gregarious behaviour in these late Campanian theropods.  

The study of new specimens from the Stephanian shales of Montceau-les-Mines confirms and enlarges the number of groups already known in this area. Among the Palaeonisciforms, “form A" is now known more completely, although no diagnosis or name can yet be given for it. “Form B" is redescribed and its relationships with “Elonichthys robisoni" are discussed. A palaeoniscid is recorded which resembles those from Bourbon l'Archambault. The paramblypteriforms occur rather frequently, but no genera can be determined. The aeduelliforms comprise some specimens close to Aeduella blainvíllei from Muse (Autun basin), and a new genus. Comparison of the latter with two fossils from Lally allows creation of two new species and a new family. This diversification of the aeduelliforms during this middle Stephanian leads to the hypothesis that the group originated at least as early the lower Stephanian. This material prooves again the characteristic endemism of this fauna, particularly of the aeduelliforms which are known only in the Massif Central where they diversified during the Permo-Carboniferous. Biogeographical consequences are discussed. 

The genus Pachynolophus, one of the poorest known of the Palaeotheriidae, includes the brachyodont forms with reduced and non-molariform premolars and with upper molars lacking a mesostyle. Quantitative characters (divers surface indications and elongation of the teeth), while demonstrating a close relationship to Hyracotherium, permit a better differentiation of the genus, confirm its specific splitting, and permit the distinction of three lineages. The skull from Memerlein is taken as the type of a new species, P. Iavocati, of which the dentition is extremely characterized by its lophiodonty, the strong reduction of the premolars and the reduction of the cingula. This characterization testifies to a late age which extends the existence of the genus quite near to the Eocene-Oligocene limit. Compared with the only two skulls known of related species (Hyracotheríum vulpiceps and Pachynolophus Iivinierensis), that from Memerlein is distinguished by progressive characters affecting diferent regions but most particularly the braincase; it is not possible, however, to isolate within this evolution the part which leads to a systematic differentiation. Modernization is translated by a considerable increase in size of the braincase, principally in the frontal region, a development of the facial region with anterior displacement of the dental series and a greater specialization of the masticatory apparatus. This evolution parallels the history of the Equidae of the North American early Tertiary, but certain particularities, the form of the alisphenoid, the presence of an anterior frontal foramen, and the structure of the paroccipital apophysis, testifies to the independance of the European forms. 

The two species of Sandgrouse from Quercy, Pterocles validus MILNE-EDWARDS and P. larvatus MILNE-EDWARDS, are ascribed to the genus Archaeoganga MOURER-CI-IAUVIRÉ which includes a third, very large species, A. pinguis. The sandgrouse of Saint-Gérand-le-Puy, Pterocles sepultus MILNE-EDWARDS, is ascribed to a new genus, Lepzoganga. This form appears in the Upper Oligocene of Quercy, in Pech Desse and Pech du Fraysse localities, and is still present in the Lower Miocene of Saint-Gérand-le-Puy and Paulhiac. Recent sandgrouse live in semidesert or desert areas. The indications provided by mammal and bird faunas in the localities where sandgrouse were found, confirm that the paleoenvironment was open and arid. The morphological study of these fossils indicates that, in the Upper Eocene, the Pteroclidae were already completely individualized with respect to Charadriiformes.
  

We report the discovery of amniotic eggshells and theropod teeth from the Late Cretaccous  period in Morocco.The megaloolithid family represents the only known dinosaur egg remains in the north part of Africa and attests indirectly to the occurrence of sauropod dinosaurs (titanosaurids) in the Maastrichtian of Africa.

  

Near "la Lieude", in the Lodève basin, more than a thousand of footprints are distributed in a twenty of trackways which amounts to 220 m length. They have been found on calcareous siltstone level in the B site named also "Réserve Naturelle Volontaire". This last is located in the Saxonian summit dated Upper Permian. "La Lieude" tracks are described by using statistical methods then they are compared with others from the world Permian. What allows to distinguish 4 following ichnotaxa: Lunaepes ollierorum nov.ichnosp., Merifontichnus thalerius nov. ichnogen. and nov. ichnosp., Planipes brachydactylus nov.ichnosp. and Brontopus circagiganteus nov. ichnosp. All these traces are attributed with possibility or probability to Therapsida or to Therosauria, except Brontopus circagiganteus nov. ichnosp. that could be due to Caseamorpha. All these animals whose sizes have been estimated between l and 5 m lived probably in a playa environment.The biological and sedimentological data from "la Lieude" footprints levels compared with informations provided by the tracks orientations, suggest the following scenario. Animals coming from the North have crossed a sandy channel bank with plants zones by directing to the South for the majority. Maybe, they were going to the lacustrine part of the playa, close to "la Lieude" footprints that they have just trampled on. 

Rodents first appear in the latest Paleocene or earliest Eocene as very fragmentary specimens (Family Paramyidae) known largely from a single locality. After this sparse beginning, rodents are usually abundant in the North American record if proper recovery methods are used. Utilization of rodents for biostratigraphic purposes depends on 1/ extinction, and 2/ replacement by evolution of endemic groups and/or incursions of Old World rodents, and rarely and late by South American kinds. These incursions are separated by relatively long periods of isolation in the Paleogene, but more episodic in the Neogene. At least 10 rodent zones can be characterized by major distinctions, and these zones can be amplified into as many as 16 with little trouble. In general, rodent genera permit as refined a zonation as do genera of large mammals. Distinction at a specific level has not been attempted herein except in the Blancan and Post-Blancan. 

Revision of the Tayassuidae from the old collections of the Quercy leads to interesting facts about taxonomic and phylogenetic problems. It could not have been well done without constant comparison with contemporaneous Tayassuidae from stratified series, as well as with European Miocene Tayassuidae and Hyotheriinae. The result is that only three species are present in the Phosphorites of Quercy: Doliochoerus quercyi FiLHOL (represented by two sub-species: the older, D. quercyi quercyi and the more recent, D. quercyi aquensis), Palaeochoerus gergorianus CROIZET et BLAINVILLE and Palaeochoerus purillus nov. sp. The two former species disappeared with Stampian time, while the third is the direct ancestor of the Aquitanian Palaeochoerus typus POMEL, which became extinct before the beginning of Burdigalian.

The comparison with Hyotheriinae brought to light new criteria for determinations, based on the study of lower premolars. These observations are condensed in a table of keys for determinations.

As a result of these studies, it came to light that the Suoidea of the Oligo-Miocene of Europe have never given rise to, except in rare cases, very long lineages. The different forms of pigs are replaced constantly in Europe by the simple game of extinctions and migrations. The reason of that discontinuity is considered here. 

The study of endocasts, or casts of the endocranial space, have played an important role in shaping understanding of mammalian, and particularly primate, brain evolution. Recently, the reconstructions of three-dimensional virtual endocasts from high-resolution computed tomography images have allowed for the visualization and quantification of endocasts in several Paleocene and Eocene primate species. Here we present the virtual endocast of MaPhQ 289 (informally known as the Montauban 9 skull), a specimen of Necrolemur antiquus Filhol 1873, a middle to late Eocene European primate of the family Microchoeridae. The virtual endocast of MaPhQ 289 reveals a lissencephalic surface morphology with expanded temporal poles and minimal overlap of the cerebellum or olfactory bulb by the cerebrum, which closely resembles the morphology of the endocast of its contemporary relative, Microchoerus erinaceus (Primates, Microchoeridae). MaPhQ 289 yields an endocranial volume (ECV) of 2.36 cm3, about 60% smaller than the volume of the most commonly cited ECV of N. antiquus. Thus, the size of the brain of N. antiquus relative to its body size is likely to be smaller than has been reported in previous literature, highlighting the importance of corroborating older ECV estimates with new evidence using 3-D imaging techniques. 

The family Didelphidae is represented by three species in the Sainte-Néboule site, phosphorites of Quercy (lower Oligocene, San Cugat's nivel): Amphiperatherium minutum (Aymard), Amphiperatherium sp. and Peratherium cuvieri (Fischer). Only the first  and  third species are abundant (88 and 97 pieces). This two populations are described. The marsupial fauna of the european lower Oligocene is not recognized in its entirety in this site. 

Pangolins have never shown a high taxic diversity and their fossil record is scarce. We report here the first discovery of a partial humerus from late Oligocene deposits in Pech du Fraysse (MP28, France). The new specimen from Pech du Fraysse is described and compared to various extant and extinct species of pangolins. It shows a suite of morphological features very similar to the humeri discovered in Saint-André (MP 26), Solnhofen (Burdigalien), and Saulcet (Aquitanian), attributed here to Necromanis franconica. The description of the specimen from Pech du Fraysse allowed us to discuss the systematics of Paleogene and Neogene pholidotans. The differences between PFY 4051 and N. franconica on the one side, and N. quercyi on the other side, might be sufficiently important to justify a generic distinction. A comparison with extant species showed that N. franconica was likely terrestrial and fossorial based on its humeral morphology. 

The entelodonts of the Quercy phosphorites are essentially known from isolated teeth. The skull remains attributed to Entelodon magnum by Kowalewsky in 1876 and the type left mandible of Enteladon depereti RÉPELIN, 1918 do not belong to this family. From a morphologic point of view, the teeth from Quercy are quite comparable to those from the type localities of E. magnum and E. deguilhemi. In the absence of distinctive anatomical criteria between E. magnum (small species from the level of Ronzon) and E. deguilhemi (large species from the level of Villebramar) they are referred, according to their size, to one or the other of these two species. However, some of them are intermediate in size. Their existence speaks in favor of a close relationship between E. magnum and E. deguilhemi and confers a real stratigraphic interest to these entelodonts of Western Europe. Finally, some anatomical characters (notch on the mesial border of P4, presence of a paraconid joined to the metaconid, trigonid higher than talonid) considered until now as peculiar to the North American Archaeotherium and not yet cited in its European contemporary Entelodon have been brought to light on the teeth from Quercy.

  

The present study of Artiodactyla from the Middle Eocene of the Geiseltal lignite beds concems systematics, biostratigraphy, and palaeoecology on the basis of 174 specimens: isolated remains to more complete skeletons. Instead of the formerly known five species of two families are now recognized 14 species of the Diacodexeidae, Dichobunidae, Cebochoeridae, and Haplobunodontidae. New species are Aumelasia maniai, Anthracobunodon neumarkensis, Masillabune franzeni. Four species of the Geiseltalfauna are definitely known from elswere, and five species are closely related to those from other European localities. Evidently the faunal situation of artiodactyls during the Middle Eocene of Europe was largely uniform. The distribution of artiodactyls within the sequence of the Geiseltal strata corroborates the biostratigraphical concept of the land mammal age Geiseltalian (Franzen & Haubold l986b) as well as the mammalian reference levels MP 11-13 (Franzen 1987). Reconstructions of the skulls and skeletons allow conclusions on the functional morphology and palaeoecology of the artiodactyls of the European Middle Eocene 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

We present here the preliminary results from a new nesting site "Sextius-Mirabeau", discovered close to the historical centre of Aix-en-Provence in Upper Cretaceous deposits. This extensive megaloolithid nesting ground, the first large scale excavation (3225 m²) in France, yielded more than 530 eggs, some organized in clusters. Their study reveals the reproductive strategy of the megaloolithid egglayer group which indicates nesting fidelity and gregarious behaviour.  

Based on isolated teeth two new scyliorhinid shark species, Scyliorhinus biformis nov. sp. and Scyliorhinus suelstorfensis nov. sp., are described from the Sülstorf Beds, early-middle Chattian, of Mecklenburg, northeastern Germany. They form part of a speciose assemblage of necto-benthic sharks and batoids which populated the warm-temperate to subtropical upper shelf sea of the south-eastern North Sea Basin. 

The dentition of selachians is characterized by an often very pronounced heterodonty involving a great morphological diversity. Despite this fact, the dentitions of selachians can be grouped in a rather reduced number of dental types corresponding to trophic adaptations: grasping, tearing, cutting, crushing, grinding and grasping-grinding type. The numerous exemples of convergence and parallelism that can be observed in fossil selachians and between Recent and fossil ones is the result of this reduced number of dental types. These dental specialisations allow to try a reconstruction of the way of life of fossil forms. 

The study of all the available skulls allows us to review the systematic relationships of the longirostrine Steneosaurus from the Middle Jurassic of western Europe. Up to now, Aalenian and Bajocian deposits have not yielded any significant Steneosaurus remain. In the Bathonian, the only valid longirostrine species, S. megistorhynchus, is known in the Britain-Normandy Basin, the Poitou and the Lorraine. In the Callovian, most of the longirostrine Steneosaurus remains can be attributed to the species S. leedsi. Nevertheless, some remains from the Middle Callovian of Poitou (France) show important differences with S. leedsi. A new Steneosaurus species, only known in Poitou, is created and named S. pictaviensis. The specific characters are carried by the skull (preorbital pit well marked, orbit and ptetygoid fossae shapes), by the mandible (symphysis shape) and by the teeth (ornamentation). S. megistorhynchus is probably situated near the stem of the Callovian species but remains from the Bathonian and Lower Callovian are very scarce and it is very difficult to precise the phylogenetic relationships between the longirostrine species of the Middle Jurassic.