A new Tertiary megadermatid is described from Rackham's Roost Site, a Pliocene limestone cave deposit on Riversleigh Station, northwestern Queensland, Australia. It appears to represent the first Australian record of Megaderma GEOFFROY, 1810, a genus otherwise known from Tertiary African and European taxa and the living Asian species M. spasma (LINNAEUS, 1758) and M. (Lyroderma) lyra PETERS, 1872. Megademza richardsi n. sp. is one of the smallest megademiatids known. It exhibits a mixture of plesiomorphic and autapomorphic features, the latter appearing to exclude it from being ancestral to any living megadermatid. The new species is one of eight megadermatids identified from the Australian fossil record, most of which are referable to Macroderma MILLER, 1906. 

A new genus, Eppsillycteris, is erected for Adapisorex? allglicus COOPER, 1932, from the earliest Eocene Blackheath Beds of Abbey Wood, London, England. Various derived character states indicate that it belongs to the order Chiroptera (bats) rather than to the extinct "insectivore" family Adapisoricidae. Other derived character states are shared with fossil and modern members of the family Emballonuridae. Placement of the new genus in this family extends the record of the Emballonuridae back in time by about 10 million years. It is the earliest record of a modern bat family and one of the earliest bats. This implies that the differentiation of at least some modern bat families took place in the Palaeocene, where no authenticated records of bats yet exist. The primitive characters of the earliest bats make the family Nyctitheriidae an unlikely stem group for the order Chiroptera. A tentative plausible alternative exists in some unnamed upper molars from the Palaeocene of Walbeck, Germany. Wyollycteris chalix, described as a bat from the Late Palaeocene of Wyoming, U,S.A., fits better in the family Nyctitheriidae. 

The discovery in the Turolian (Late Miocene) of Dytiko 3 (Macedonia, Greece) of a complete skull with mandibles and cervical vertebrae, atlas and epistropheus, is a very important contribution to the knowledge of the subfarnily Chalicotheríinae. After the description, the comparison with other specimens of Miocene chalicotheres permits the revival of the generic name Macrotherium with a new species M. macedonicum. This genus is mainly characterized by a short snout and an inflated cerebral skull. It coexists during the Miocene with Chalicotherium. A cladistic analysis leads to conclusion that the species which has been described from the Early Middle Miocene of Rusinga must be identified as the type-species of a new genus: Butleria.





  

The Late Triassic locality of Saint-Nicolas-de-Port (Meurthe-et-Moselle, France) has yielded numerous isolated teeth belonging to archosauriform reptiles. The following tooth groups can be identified: heterodont phytosaurs, the pterosaur Eudimorphodon, the prosauropod dinosaur Plateosaurus, three types of putative ornithischian teeth and 13 types of carnivorous Archosauriformes indet. Apparent venom-conducting teeth belonging to a new taxon of ?Archosauriformes (Graoullyodon hacheti nov. gen. nov. sp.) are also described. From a palaeogeographical point of view, the ornithischian teeth from Saint-Nicolas-de-Port (if their attribution is confirmed) are the oldest fossils of this group in Europe. The biostratigraphic distribution of the tooth forms mostly suggests a Late Norian or Early Rhaetian (depending on current interpretations) age of the deposits, but do not provide more precisions than fossils previously described from the area. The dietary habits and, consequently, the palaeoecological relationships of the different vertebrate groups discovered at Saint-Nicolas-de-Port are tentatively established: the ornithischian and prosauropod teeth reflect a herbivorous diet, whereas the other archosauriform teeth are probably from camivores or omnivores.

  

Revision of the hipparion material from Montredon, including newly excavated and other unpublished specimens brings evidence of specific heterogeneity.
A fragmentary very small MC III seems very close to H. macedonicum from the upper Vallesian of Ravin de la Pluie, Greece. In that same site was also found a large hipparion.
Most of the Montredon material is referred to H. depereti. This species associates characters usually found in Vallesian hipparions (highly plicated upper cheek teeth, deep vestíbular grooves on the lower cheek teeth, robust metapodials) with characters more frequent in Turolian forms (middle size, lack of confluence in the upper premolar fossettes, lack of ectostylids on the adult lower cheek teeth, well developped keel on the MC III, facette for the 2nd cuneiform present on all MT III). H. depereti shares some of these characters with the Spanish and Portuguese hipparions transítional between the Vallesian and the Turolian (Masia del Barbo, Azambujeira) but is not identícal to any of them. The upper Vallesian hipparion material from Diavata, Greece, probably belongs to H. depereti but not the large hipparion rests from Ravin de la Pluie.
Thus, Montredon and Ravin de la Pluie may well share the same small species, H. macedonicum, but they differ in the associated one: middle-sized H. depereti at Montredon, large-sized H. primigenium at Ravin de la Pluie. Both sites, however, give evidence of small hipparions during the Vallesian, coexisting with other larger species.
The present paper also proposes an adaptation of the Kiesewalter's indices that calculates the height at the withers using the metapodial length, and discusses indices proposed by Gromova and by Sen et al. to express the relative development of the metapodial keel. 

New Dendromus species (Rodentia, Muroídea) from Langebaanweg (Pliocene, South Africa). Stratigraphical and paleoecological consequences.

Two new species of Dendromus are described from the Langebaanweg site which precises the evolutionary trend among this genus in South Africa and gives further paleoenvironmental indications. Two evolutionary stages are described: D. darti nov. sp. shows low-crowned molars with bunodont cusps and its more closest relative would be D. melanozis from the Cape region. On the contrary, D. averyi nov. sp. is more lophodont and is better related with the modem D. melanotis. Both species are at a less evolved stage than the Dendromus sp. from Laetolil Beds at Laetoli. The Langebaanweg deposits cannot still be dated by biostratigraphy but they clearly cannot be older than the basis of Pliocene times. The association of Dendromus and Mystromys in the same levels indicates a grassland environment with woodland patches as well as probable swamps. 

The St-Dézéry local fauna (3 reptile-, 4 mammal species) is approximately of the same age as the La Débruge or the Ste-Néboule faunas. It conduces to a better dating of the limestones underlying the Célas sandstones. A large part of a mandible of Amphimeryx was found there, which documents the record of this family of small artiodactyls 

The present study of Artiodactyla from the Middle Eocene of the Geiseltal lignite beds concems systematics, biostratigraphy, and palaeoecology on the basis of 174 specimens: isolated remains to more complete skeletons. Instead of the formerly known five species of two families are now recognized 14 species of the Diacodexeidae, Dichobunidae, Cebochoeridae, and Haplobunodontidae. New species are Aumelasia maniai, Anthracobunodon neumarkensis, Masillabune franzeni. Four species of the Geiseltalfauna are definitely known from elswere, and five species are closely related to those from other European localities. Evidently the faunal situation of artiodactyls during the Middle Eocene of Europe was largely uniform. The distribution of artiodactyls within the sequence of the Geiseltal strata corroborates the biostratigraphical concept of the land mammal age Geiseltalian (Franzen & Haubold l986b) as well as the mammalian reference levels MP 11-13 (Franzen 1987). Reconstructions of the skulls and skeletons allow conclusions on the functional morphology and palaeoecology of the artiodactyls of the European Middle Eocene 

The numerical importance of the Condylarths in the Cernaysian fauna is discussed. The Condylarth family, Tricuspiodontidae, is reviewed in the light of new material and its close relationships to the Phenacodontidae is suggested ; one new species is recognized : Tricuspiodon sobrinus. European Arctocyonidae are reviewed and the recentclassification of Van Valen is briefly commented on. Also,  the arctocyonine Landenodon is described for the first time in Thanetian (Late Paleocene) sediments ; two new species are proposed : T. lavocati and T. phelizoni. 

The wear facets of primitive rodents can be homologized with those of primitive primates and ungulates. As in primates, the jaw movement was ectental, with an increased anterior component in the lingual phase (phase ll). The buccal phase (phase I) in rodents approaches the horizontal and it tends to be reduced in importance in comparison with the lingual phase. ln more advanced rodents the efficiency of grinding is increased by the development of additional cutting edges of enamel (e.g. enlargement of hypocone, development of mesoloph and lingual sinus). The buccal phase movement becomes lined up with the lingual phase movement to form a single oblique chewing stroke,resulting in planation of the crown. As the stroke becomes more longitudinal (propalinal) the enamel edges become more transverse. In Muridae propalinal chewing evolved before the loss of cusps, facets were reorientated and additional cusps developed. 

The complex taxonomic history of the paramyid rodent genus Ailuravus is reviewed. It has been described as Hyracotherium, as a creodont carnivore and as a lemuroid primate - errors at the ordínal level that are most unusual for a rodent. The genus is a member of the poorly known subfamily Ailuravinae, probably derived from some European Early Eocene species of Paramys. Aíluravus was a large arboreal paramyid with highly rugose cheek teeth, very well developed hypocone, and a remarkably weak lower incisor. It was tropical to subtropical. Three named species are recognized, A. macrurus from the Lutetian of Messel; the genotype, A. picteti, from Egerkingen, Buchsweiler and the Geiseltal, slightly later in the Lutetian; and A. stehlinschaubi, new name, from the Bartonian of Mormont-Eclépens and Robiac. One or more unnamed species are present in the Ypresian of Cuis. The species are close to a phyletic sequence. No later representatives of the genus are known. The late Eocene to earliest Oligocene North American paramyid Mytonomys, whose relationships have been obscure, is tentatively referred to the Ailuravinae. 

An unpublished mandible of the large hyracoid Titanohyrax andrewsi from the early Oligocene Jebel Qatrani Formation, Fayum Depression, Egypt is described. This specimen has a twofold importance. Firstly, it opens an unexpected window on early paleontological research in the Fayum because it was discovered as early as 1904 by the French paleontologist René Fourtau during an expedition to the Fayum organized by the Muséum National d’Histoire Naturelle, Paris (MNHN). This expedition has remarkably never been mentioned in the literature. Secondly, the mandible documents the best-preserved specimen of T. andrewsi, permitting a revision of one of the very rare Paleogene hyracoids. Interestingly, the new mandible was discovered two years before the first report of the species by Charles W. Andrews. The hypodigm of T. andrewsi is reviewed and the dentition as a whole is compared in detail, notably with other Titanohyrax species from the Fayum. The validity of the large Titanohyrax “schlosseri” species is discussed, but a pronounced sexual size dimorphism for T. andrewsi is favoured. 

Anthracotheres from the Pondaung Formation, Myanmar, are considered as one of the most primitive artiodactyl groups and they represent the oldest known record in the world. Thus, the understanding of this group has numerous implications for evolutionary biology and biochronological correlations. However, the systematlcs of these mammals has been interpreted in different ways, and the main debate focuses on the number of taxa represented in the Pondaung Formation. The revised taxonomy proposed here is mainly based on the relative development of the upper molar W-shaped ectoloph, system of crests and stylar cusps, and on body size. On the basis of these characters, they are classified into four genera including six different species. Two well-known genera, Anthracotherium and Anthracokeryx, are validated and more precisely diagnosed. Anthracokeryx possesses a better developed W-shaped ectoloph, system of crests and stylar cusps than Anthracotherium, which displays notable differences with the more derived representatives of this genus. Both of these Pondaung genera show evidence for sexual dimorphism. However, the incompleteness of fossil material fueled a debate concerning the status of two additional Pondaung anthracotheres, Siamotherium and Anthracohyus. The latter genus is of uncertain affinities, but it has been considered as a hippopotamid ancestor. Despite new material attributed to these two forms, additional discoveries are still required to establish their taxonomic status. The hypothesis that Southeast Asia was the centre of origin of Anthracotheriidae is supported by the retention of numerous primitive dental characters in these taxa and by the antiquity of the Pondaung Formation, to which an age of 37 My is now generally accepted. 

Morphological descriptions are given of Eocene mammals from the locality Toru Ajgyr (NEKyrgyzstan) that were excavated in 1997 and 1998 in a cooperation between the Martin-Luther-University Halle (Germany), the Zoological Institute in St. Petersburg (Russia) and the Seismological Institute in Bishkek (Kyrgyzstan). The species found belong mostly to perissodactyls, as Lophialetes sp., Teleolophus sp. and brontotheres. The primitive ungulate family Olseniidae is represented by a complete foot skeleton of cf. Olsenia sp. In addition, postcranial materials of Gobiatherium mirificum (Dinocerata) and of artiodactyls have been collected and are described herein. Based on mammals, the locality is part of the Asian Land Mammal Age Arshantan and is stratigraphically equivalent with the Bridgerian Land Mammal Age in North America and with the lower and middle Geiseltalian of the European Middle Eocene. 

Cranial anatomy of a late Miocene rhizomyid, Brafhyrhizomys cf. B. pilgrimi, provides new evidence on the origin of the dorsal, round infraorbital foramen of living rhizomyines. Primitive rhizomyids retain a myomorphous keyhole foramen with a long ventral slit that retracts upward during the evolutionary history of the Rhizomyidae. The primitive condition of the elongated ventral slit is represented by Kanisamys sivalensis. Among later burrowers the foramen shows progressive dorsal migration, the ventral slit terminating midway up the snout in B.tertracharax and B. choristos ; well above the midline of the snout in Brachyrhizamys cf. B. pilgrimi. Apparently this transformation began earlier among Rhizomyinae than among Tachyoryctinae and continued to a more derived stage in rhizomyines. ln living Rhizomyx the ventral slit is absent and only a high round hole remains at the anterior end of the zygomatic arch. 

 Detailed anatomical description of arsinoithere cranial remains from the Lower Oligocene, Fayum Depression, Egypt, provides the basic data for a systematic investigation. All cranial and some postcranial features are assessed from a phylogenetic standpoint. Several soft tissue characters are then added to a cladistic analysis based on 54 derived ungulate morphological characters. The resulting phylogenetic hypothesis implies that perissodactyls, sirenians, proboscideans and arsinoitheres constitute a monophyletic unit (5 synapomorphies). However, increasing the tree length by 3 steps reveals a closer association between hyraxes and perissodactyls. Nevertheless, 13 synapomorphies link proboscideans, sirenians and  arsinoitheres to the exclusion of all other ungulates. Form of the sphenopalatine and ethmoid foramina, recurved posttympanic process, absence of a fenestra rotundum in the petrosal, vestigial paroccipital process of the exoccipital and the highly unusual absence of a hypoglossal foramen in the skull, imply a robust sister-group relationship between arsinoitheres and proboscideans. In this analysis artiodactyls share only one derived character with all other ungulates studied. Monophyly of Ungulata, including Artiodactyla, is therefore only weakly supported. It is argued that pedal anatomy of hyraxes is non-homologous with that of Tethytheria. Arsinoitherium should now be classified within Tethytheria, sharing a sister-group relationship with Proboscidea. Hyraxes are excluded, thus refuting the concept of Paenungulata. However, monophyly of the wider concept, Pantomesaxonia, containing hyraxes, perissodactyls, sirenians, proboscideans and now, arsinoitheres, is supported by this study. 

The rodent faunas of five afghan localities found in 1976 and 1977 (Sherullah, Ghazgay, Pul-e Charkhi, Dawrankhel 14 and 15) are studied.
The rodents (Muridae, Cricetidae and Rhizomyidae) represent 8 genera and 10 species. The detailed description of the 2 new genera and 7 species diagnosed in 1979 is given. An other species is created : Pseudomeriones crapouilloti n. sp. These faunas precise the origin and diversification of Muridae and Cricetidae. A phyletic lineage known in Afghanistan is represented in East Africa by a ramus or a collateral lineage. The five localities are dated from Lower Turolian to Ruscinian. They constitute the frame of a chronologie scale for the Upper Continental Neogene of Afghanistan.
The study of afghan material brings new data to the biogeography of Old Word's rodents during the Upper Neogene; from Pakistan to Europe and Africa, a rather warm and damp province would have existed till Upper Miocene; after which (in the mio-pliocene epoch) it would have divided into 3 parts, by aridification of the central area. 

The Condylarths from Hainin (Hainault, Belgium) show no affinity at the generic level to those known in other Paleocene localities of Europe and North America ; they are described as new forms : Monshyus praevius n. gen., n. sp. and Prolatidens waudruae n. gen., n. sp. Monshyus praevius, discovered in only one of the levels in the excavation at Hainin, is similar to the genera Microhyus TEILHARD and Louisina RUSSELL ; with them it is included in the subfamily Louisininae (Hyopsodontidae). With respect to Microhyus and Louisina, Monshyus is distinguished by the precociously modern aspect of its upper molars, the only teeth that are referable. Prolatidens waudruae, known only by lower molars, was found in several levels in the pit at Hainin. It is an arctocyonid presenting possible relationships to the North American form Oxyprimus galadrielae ; it therefore has been provisionally attributed to the subfamily Oxyclaeninae. If this attribution is confirmed, this species will constitute the first and only representative of the group in Europe. 

Based on material found in about 15 localities the relationships of the genera Microparamys and Glirarus have been studied. One new genus, two subgenera and three species [Microparamys (Sparnacomys) chandoni n. subgen. and n. sp., Microparamys (Pantrogna) russelli n. subgen., Eoglirarus wildi n. gen. and n. sp., Gliravus meridionalis n. sp.] as well as the publication
of numerous new facts concerning species previously reported, support the phyletic scheme proposed. The latter shows that the origin of the Gliravinae is to be sought among the very small and still rather poorly known Microparamys species of the early Eocene. Gliravus: robiacensis can be considered as the common ancestor to different lineages not only of Glirarus but also of modern genera (Peridyromys, Glirudinus and Microdyromyx), at the origin of which Peridyromys micio, although difficult  to interpret, occupies a similar place.

The stratigraphic conclusions permit more detail in the chronologie succession of the localities studied. The paleoecologic and biogeographic aspects lead one to the problem of the oligocene "Grande Coupure" through the study of this group.

  

New records of terrestrial mammals from the Qasr el Sagha Formation, Fayum Depression, Egypt are reported, and the stratigraphic occurrences of these fossils noted. These include additional specimens of Moeritheríum, Barytherium, and anthracotheres, as well as the oldest record of a hyracoid in the Fayum.These Eocene mammals occur almost exclusively in the alluvial deposits of the Dir Abu Lifa Member of the Qasr el Sagha Formation and show close affinities to the faunas from the lower sequence of the Jebel Qatrani Formation. There is no evidence of a more marked faunal discontinuity between the Qasr el Sagha and Jebel Qatrani Formations than there is across any of the three major breaks in sedimentation that exist within the Jebel Qatrani Formation. The faunal similarities between fossils of the lower sequence of the Jebel Qatrani Formation and of the upper part of the Qasr el Sagha Formation is consistent with recent paleomagnetic dating that suggests that these rocks differ in age by only one to two million years.