Nyctereutes megamastoides (Pomel) from the Villafranchian of the Auvergne and from Saint-Vallier presents cranial and dental characters sufficiently close to those of the late Pliocene canid from Perpignan (Roussillon), described by Depéret under the specific name of Canis donnezani belonging to the same genus Nyctereutes. The extinction of the European "Nyctereutes" group seems due to the too great alimentary specialization of this canid, whereas the Asiatic lineage represented in the Villafranchian by Nyctereutes sinensis Schlosser and at present by Nyctereutes procyonider Gray was able to maintain itself probably by means of a profound change in its alimentary regime. 

We report the discovery of amniotic eggshells and theropod teeth from the Late Cretaccous  period in Morocco.The megaloolithid family represents the only known dinosaur egg remains in the north part of Africa and attests indirectly to the occurrence of sauropod dinosaurs (titanosaurids) in the Maastrichtian of Africa.

  

A new genus and species of haplobunodontid artiodactyl, Hallebune krumbiegeli nov. gen. nov. sp., is described from the middle Eocene Geiseltal Fauna (Saxon-Anhalt, Germany). Its stratigraphic range is restricted to the MP 13 ("obere Mittelkohle") of the Geiseltalian. The material consists of seven fragments, representing both upper and lower jaws. It is indicated, that H. krumbiegeli was a small bunodont herbivore. The size and primitive character of the teeth suggest that the genus possibly is directly derived from Diacodexeidae.The new material forms the basis for reinterpreting the haplobunodontid phylogeny. 

In the course of a revision of the Equoidea numerous dentitions as well as a partial skeleton of a Phenaeodont were discovered from the Middle Eocene lignite beds of the Geiseltal locality. These fossils are recognized as a new genus and species of Phenacodontidae : HaIlensia matthesi n.g. n.sp.. The species is present in the « untere und obere Unterkohle ›› (uUK, oUK = the lower and upper part of the Lower Coal Seam) as well as in the « obere Mittelkohle ›› (oMK = the upper part of the Middle Coal Seam). Two fragmentary upper jaws described and figured by Matthes (1977) as Propachynolophus gaudryi are also belonging to Hallensia matthesi. Thus the decisive argument for classifying the " Unterkohle " of the Geiseltal section as Lower Eocene has to be dropped. Another relict form of the Geiseltal is Esthonyx tardus n. sp. documented by a fragmentary mandible coming from the « untere Unterkohle ››. This is the latest Tillodont from Europe. Contrasting to E. munieri from the european Lower Eocene the dentition of E. tardus is morphologically more progressive. 

This paper is about the oldest African rodents faunas, from the late Early Eocene, or early Middle Eocene, Glib Zegdou (Algeria) and Chambi (Tunisia) localities. Five species are described and figured, belonging to a new family here created, the Zegdoumyidae.

This family is compared to the Early and Middle Eocene rodents families from Asia, Europe and North America (Chapattimyidae, Yuomyidae, Gliridae, Theridomyidae, lschyromyidae and Sciuravidae), as well as to those known from the Late Eocene African locality Bir El Ater (Anomaluridae and Phiomyidae).

On the one hand, it seems clear that the African endemic Anomaluridae arise from the Zegdoumyidae. On the other hand, the lschyromyidae, or primitive Sciuravidae, may be the most reliable ancestral groups for the Zegdoumyidae. Thus, this new family can be considered as the sister group for the American Sciuravidae on the one hand, and for the European Gliridae on the other hand.

The biogeographical consequences of these phylogenetic hypotheses are discussed. A new phase of communication between Europe and North Africa is inferred, during the Early Eocene. It has been followed by a short period of endemism, allowing the adaptive radiation for the Zegdoumyidae, preceding the immigration of the Phiomyidae, during the Late Eocene, probably from Asian relatives. 

The Laprade fauna is chronologically situated between those from Egerkingen and Lissieu and consequently, is close to the MP 14 reference-level of the European mammalian biochronological scale (Symposium of Mainz, 1987).
This new fauna is presently the oldest known in the Quercy phosphorites, formerly the Le Bretou fauna (MP 16) was considered as the oldest one. The Laprade fauna includes 21 species which belong in 7 mammalian orders (Marsupialia: Amphiperatherium bastbergense, Amphiperatherium goethei; Apatotheria: Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp.; Lipotyphla: Saturninia cf. mamertensis, Saturninia cf. intermedia; Chiroptera: Vespertiliavus lapradei nov. sp.; Rodentia: Protadelomys cf. lugdunensis, Elfomys nov. sp.; Primates: Nannopithex cf. filholi, cf. Pseudoloris or Pivetonia; Perissodactyla: ?Palaeotherium ?castrense, small-sized Palaeotherium sp., Plagiolophus sp., Anchilophus sp.; Artiodactyla: Dichobune cf. robertiana, Mouillacitherium cartieri, Tapirulus cf. depereti, Mixtotherium priscum, Pseudamphimeryx schlosseri, and Artiodactyla indet). Sixteen of these species are mentioned for the first time from the Quercy faunas.
The recognition of a new apatemyid, Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp., attests to an early origin of a lineage known in the Late Eocene. The emballonurid bat Vespertiliavus lapradei nov. sp. is presently the earliest record of this genus and family. This Auversian fauna leads to discuss the age of taxa showing archaic features. These taxa were defined on specimens collected in the Quercy during the last century and have never been found in the Quercy localities recently investigated. This fauna bears also evidence of a karstic filling episode older than those previously dated by fossils in the Quercy Jurassic. 

A classification should be both usable and useful,not too complex either in the amount of splitting or in the number of hierarchies involved, and not so simple as to give a false assurance of knowledge of relationships. Classifications are only possible because we do not have complete knowledge of the evolution of the organisms concerned because gaps in the record are necessary to allow the separation of the various taxa. Rodent classification is complicated by the large number of organisms involved and by the geat amount of parallelism that has taken place in the evolution of any and all features. If several independent features are characteristic of a certain taxon, should an effort be made to define the group on the basis of all the features, or should only one be selected as the determinant ? Unless the evolution of the several features was closely linked, the former solution will sooner or later lead to insurmountable problems. 

Le Quercy est aujourd 'hui un vaste plateau calcaire, parcouru par un réseau karstique actif, profondément
entaillé par des vallées aux falaises abruptes, comme celles du Lot ou du Célé. Sur un sol peu épais domine la forêt de chênes, accompagnés de cornouillers, érables, genévriers. La faune est pauvre, peu diverse, et les nombreux chasseurs se satisfont de gibier d'élevage ...

Il y a trente cinq millions d'années environ, le paysage était bien différent. La période de l'Eocène supérieur, qui s'achevait, avait été chaude et humide, si l'on se réfère à la fois aux paléotempératures (calculées à partir de sédiments marins extra·européens) et aux restes fossilisés de végétaux typiquement tropicaux. 

Bats are among the most numerous and widespread mammals today, but their fossil record is comparatively meagre and their early evolution poorly understood. Here we describe a new fossil bat from dental remains recovered from late Early Eocene sediments at Prémontré, northern France. This 50 million-year-old bat exhibits a mosaic of plesiomorphic and apomorphic dental features, including the presence of three lower premolars, a single-rooted p3, short p4 with metaconid, myotodont lower molars and a tall coronoid process of the dentary. This combination of features suggests it is an early member of Vespertilionidae, today’s most speciose and geographically widespread bat family. The Prémontré bat has bearing on hypotheses about the origins of vesper or evening bats (Family Vespertilionidae), as well as crown-group chiropterans.


  

Résumé :
L’étude des rongeurs Theridomorpha permet de suivre le déroulement d’une radiation adaptative pendant toute sa durée (Eocène moyen-Oligocène terminal), sur un territoire restreint à l’extrémité ouest de l’Europe Occidentale. Dans ce papier, la phylogénie de ce groupe est établie à partir d’une analyse cladistique reposant sur l’examen de 315 caractères (310 dentaires). Le groupe d’intérêt comprend 110 des 132 espèces (24 genres) de Theridomyoidea et deux genres encore inclus jusqu’ici dans les Reithroparamyinae qui rejoignent les Theridomorpha. Les groupes externes comprennent des Glires basaux, Cocomys, Tanquammys et 16 Ischyromyiformes. Un cadre phylogénétique robuste est produit, qui permet de clarifier la systématique des Theridomorpha. La position des Remyoidea (nov. sup.fam.) au sein des Ischyromyiformes, extérieure aux Theridomorpha, est confortée. Les Protadelomys et Tardenomys sont à la base des Theridomyoidea, avant la séparation en deux clades correspondant aux familles Pseudosciuridae et Theridomyidae. Les sous-familles sont consolidées : Pseudosciurinae et Sciuroidinae pour les Pseudosciuridae ; Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, auxquelles s’ajoute au moins une nouvelle sous-famille (Patriotheridomyinae), pour les Theridomyidae. La topologie des chrono-espèces (sensu Simpson), traitées antérieurement comme lignées évolutives, apparaît dans la plupart des cas sous forme de clades successifs dans lesquels les espèces sont le plus souvent arrangées de manière pectinée, émergeant dans l’ordre stratigraphique. L’analyse des caractères aux principaux nœuds permet de consolider les caractères diagnostiques des taxons et les tendances évolutives, ainsi que de discuter des divers parallélismes et convergences dans l’évolution des structures et patrons dentaires (e.g., émail des incisives unisérié chez les Issiodoromyinae et les Patriotheridomyinae, ou pseudo-multisérié chez les Blainvillimys les plus hypsodontes, les Protechimys et Archaeomys ; patrons dentaires téniodontes ; allongement des dents déciduales chez les Patriotheridomyinae, Issiodoromyinae et Theridomyidae ; sélénodontie ou lophodontie). Les dynamiques évolutives traduites par les changements morphologiques sont mises en relation avec les variations environnementales. Enfin, les implications biochronologiques de l’évolution des Theridomyoidea sont discutées.
Abstract:
The adaptive radiation of the rodents Theridomorpha occurred during a limited time window (middle Eocene to late Oligocene), on an area restricted to Western Europe. In this paper, the phylogeny of this group is established via a cladistic assessment of 315 morphological characters (310 dental). The group of interest encompasses 110 upon 132 species (24 genera) of Theridomyoidea, and two genera formerly included within the Reithroparamyinae, and which are included here within the Theridomorpha. The outgroups include basal Glires, Cocomys, Tanquammys and 16 Ischyromyiformes. A robust phylogenetic frame is produced, which allows clarifying the systematics of the Theridomorpha. Within the Ischyromyiformes, the Remyoidea (nov. supfam.) are set apart from the Theridomorpha. Protadelomys and Tardenomys represent the earliest offshoots of the Theridomyoidea, before the dichotomy between Pseudosciuridae and Theridomyidae. It supports the former subfamilies Pseudosciurinae and Sciuroidinae within the Pseudosciuridae; and for the Theridomyidae: the Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, with at least one new subfamily (Patriotheridomyinae). The topologies of the chronospecies (sensu Simpson), formerly considered as evolutionary lineages, appear in most cases as successive clades, in which the species are generally pectinately arranged and emerging in the stratigraphic order. The analysis of characters supporting the main nodes allow consolidating the diagnosic characters of the taxa and their evolutionary trends, as well as discussing the various cases of parallelism and convergence in the evolution of structures and dental patterns (e.g., uniserial incisor enamel for Issiodoromyinae and Patriotheridomyinae, or pseudo-multiserial for the most hypsodont Blainvillimys, Protechimys and Archaeomys; taeniodont dental patterns; lengthening of deciduous premolars for Patriotheridomyinae, Issiodoromyinae and Theridomyidae; selenodonty or lophodonty).
Evolutionary dynamics are analysed with respect to environmental changes. Finally, biochronological implications of the evolution of Theridomyoidea are discussed.
  

Seventeen species of cricetid rodent are recognized and described from lower and middle Siwalik deposits in the Potwar Plateau of Pakistan. These species are grouped in three categories, characterized as subfamilies (e. g., Megacricetodontinae, Myocricetodontinae, and Dendromurinae); an additional and more abundant category of rodents from these deposits, the Democricetodontinae, is excluded from this study, and will be described in a later study. Fifteen of the species are new, and four new genera are described. The Siwalik cricetid taxa are : Megacricetodon aquilari, n. sp.; Megacricetodon sivalensis, n. sp.; Megacricetodon daamsi, n. sp.; Megacricetodon mythikos, n. sp.; Punjabemys downsi, n. gen. & n. sp.; Punjabemys leptos, n. gen. & n. sp.; Punjabemys mikros, n. gen. & n. sp.; Myocricetodon sivalensis, n. sp.;  Myocricetodon sp.; Dakkamyoides lavocati, n. gen. & n. sp.; Dakkamyoides perplexus, n. gen. & n. sp.; Dakkamys asiaticus, n. sp.; Dakkamys barryi, n. sp.; Dakkamys sp.; Paradakkamys chinjiensis, n. gen. & n. sp.; Potwarmus primitivus, n. gen.; and Potwarmus minimus, n. gen. & n. sp. This diverse record of middle Miocene small mammals illuminates a profound radiation of cricetid rodents in southem Asia, the effects of which were felt in Europe and Africa as well as the rest of Asia. 

The description of original material of anthracothere and proboscidean in the new locality of Bir el Ater 3 from East Algeria, and a thorough review of early Libycosaurus remains of Bir el Ater 2 allows us validating L. algeriensis as the smallest and earliest species of Libycosaurus and probably the earliest migrant of the genus from Asia. The presence of a Tetralophodon in the Neogene Nementcha formation might represent the earliest occurrence of the genus in Africa. These original fossil remains allow us to discuss the age of the Neogene part of the Nementcha formation close to the Serravalian/Tortonian boundary. 

Based on material found in about 15 localities the relationships of the genera Microparamys and Glirarus have been studied. One new genus, two subgenera and three species [Microparamys (Sparnacomys) chandoni n. subgen. and n. sp., Microparamys (Pantrogna) russelli n. subgen., Eoglirarus wildi n. gen. and n. sp., Gliravus meridionalis n. sp.] as well as the publication
of numerous new facts concerning species previously reported, support the phyletic scheme proposed. The latter shows that the origin of the Gliravinae is to be sought among the very small and still rather poorly known Microparamys species of the early Eocene. Gliravus: robiacensis can be considered as the common ancestor to different lineages not only of Glirarus but also of modern genera (Peridyromys, Glirudinus and Microdyromyx), at the origin of which Peridyromys micio, although difficult  to interpret, occupies a similar place.

The stratigraphic conclusions permit more detail in the chronologie succession of the localities studied. The paleoecologic and biogeographic aspects lead one to the problem of the oligocene "Grande Coupure" through the study of this group.

  

The Palaeobatrachidae until now have not been reported from France. The present note makes known the discovery of very significant remains in three Tertiary localities of France: Cernay (late Paleocene), Laugnac (late Aquitanian) and Sansan (middle Helvetian). The osteologic characters, as well as the geographic and stratigraphic distribution of the family are re-stated with added details. The Palaeobatrachidae are a European family known from the late Jurassic of Catalonia (publication in course by J. Seiffert) to the Plio-Pleistocene (pre-Mindel) of Poland and Rumania. By their way of life and certain osteologic characters they are similar to the Pipidae, but they are distinguished by profound differences. It would be rash to include them in the Pipoidea or even in the Aglossa. They are apparently an independant group, essentially holarctic, which held in Europe a role equivalent to that played by the Pipidae south of Tethys. 

The object of this work is the study of the artiodactyls of the karstic locality of Lissieu near Lyon. Among the sixteen species counted, five (Hyperdichobune nobilis, Hyperdichobune sp., Haplabunodon mulleri, Tapirulus depereti and Pseudamphimeryx schlosseri) are cited for the first time in this fauna. The description of part of the anterior dentition of «Dichobune» langi permits the referal of this species to the genus Hyperdichobune. The new selenodont species Dichodon lugdunensis  described in this work belongs, with its ancestor Dichodon simplex, to a lineage distinct from that of Dichodan cartieri and Dichodon ruetimeyeri. Finally, the taxonomic reality of the Lissieu species, Lophiobunodon rhodanicum Deperet, is justified. All the other species have been compared with those of Egerkingen. Even though with most of them no difference in age can be shown between the two deposits, some however indicate clearly by their degree of evolution that Lissieu is slightly more recent than Egerkingen. ln the history of Eocene mammals, the Lissieu fauna, because of its homogeneity, can be considered as a sure reference fauna of the Auversian period. 

An upper giraffid premolar without any indication about its origin is preserved at the Montpellier University among numerous fossils from the ruscinian formation of Montpellier. It can be related to Samotherium, of the Upper Miocene in Eastern Europe, North Africa and Asia, or more probably to Bramatherium  or Hydaspitherium of the Pliocene of South East Asia. The sedimentological study of the matrix shows a calcareous background, which may indicate that this tooth does not come from the Montpellier formation. 

The genus Microstonyx was found only in the north eastern part of Spain : M. antiquus, referable to the
Eppelsheim type, from the pyrenean basins ; M. antiquus, more progresive, from the Vallès Basin : M. major-erymanthius group, from the Teruel-Ademuz Area and from Pieru (Penedès Basin).

  

Les Carnivores Fissipèdes de Sainte-Néboule appartiennent tous au genre Cynodictis et semblent constituer une population homogène. Celle-ci se distingue suffisamment des espèces déjà décrites pour constituer un taxon particulier : Cynodictis lacustris neboulensis n. s. sp. . L'étude des variations à l'intérieur de cette population nous a conduit à reconsidérer les critères utilisés pour définir les espèces existantes et à regrouper certaines d'entre elles. Il semble qu'il demeure cependant trois lignées distinctes dans le genre Cynodictis mais le matériel nous paraît encore insuffisant pour traduire cette remarque en termes de systématique. 

Morphological descriptions are given of Eocene mammals from the locality Toru Ajgyr (NEKyrgyzstan) that were excavated in 1997 and 1998 in a cooperation between the Martin-Luther-University Halle (Germany), the Zoological Institute in St. Petersburg (Russia) and the Seismological Institute in Bishkek (Kyrgyzstan). The species found belong mostly to perissodactyls, as Lophialetes sp., Teleolophus sp. and brontotheres. The primitive ungulate family Olseniidae is represented by a complete foot skeleton of cf. Olsenia sp. In addition, postcranial materials of Gobiatherium mirificum (Dinocerata) and of artiodactyls have been collected and are described herein. Based on mammals, the locality is part of the Asian Land Mammal Age Arshantan and is stratigraphically equivalent with the Bridgerian Land Mammal Age in North America and with the lower and middle Geiseltalian of the European Middle Eocene. 

No abstract available