The first good skeletal remains of Homogalax protapirinus from the Wasatchian of the Bighorn Basin, Wyoming, indicate that this primitive tapiromorph was more plesiomorphic in many features than primitive equoids including Hyracotherium. Compared to Hyracotherium, Homogalax more closely resembles Phenacodonta (the closest outgroup of Perissodactyla for which postcrania are known) in various details of articular surfaces, muscle attachments, and proportions of the humerus, manus, and pes.Among known taxa, Homogalax most nearly approximates the plesiomorphic postcranial skeletal anatomy of Perissodactyla. 

Numerous “Quaternary” large-mammal fossils have been collected since the 1830s along the Le Riège stream, near Pézenas (Southern France). More than 120 specimens are stored in the collections of the Université de Montpellier (UM) under the name “Le Riège”. A major operation aiming at relocating the palaeontological collections of the University has made it possible to group together all the specimens of interest and launch their systematic revision for the first time. The fossils belong to the Reboul (1839; 51 samples) and de Christol (1865; 18 samples) Collections and 17 samples compose the Crochet & Ivorra Collection (1998). The remaining 38 samples have no mention about the exact time and location of their finding. We provide a critical inventory with literal transcription of inscriptions on specimens and historical labels. This revision confirms the presence of two distinct faunal assemblages under the name of “Le Riège”: Saint-Palais (Early Pliocene, MN14–15) and Le Riège sensu stricto (late Early Pleistocene, most likely MNQ19). The former assemblage, with coastal affinities, is composed of the ruminants Alephis sp. and Procapreolus cf. pyrenaicus, the rhinocerotid Pliorhinus megarhinus, the gomphotheriid Anancus arvernensis and marine mammals, all emblematic taxa for the Early Pliocene of Montpellier and Perpignan. The latter assemblage documents a late Early Pleistocene fluvio-volcanic sequence, yielding the bovid Bison (Eobison) spp., the cervid Eucladoceros cf. giulii, the hippopotamid Hippopotamus antiquus, the rhinocerotid Stephanorhinus etruscus, the equid Equus cf. altidens, and the elephantid Mammuthus cf. meridionalis, plus a few specimens of uncertain taxonomic affinities. This revision underscores the interest of revisiting historical collections and further provides a starting point for future research.
  

The earliest cetaceans are found in the early Eocene of Indo-Pakistan. By the late middle to late Eocene, the group colonized most oceans of the planet. This late Eocene worldwide distribution clearly indicates that their dispersal took place during the middle Eocene (Lutetian). We report here the first discovery of a protocetid fossil from middle Eocene deposits of Senegal (West Africa). The Lutetian cetacean specimen from Senegal is a partial left innominate. Its overall form and proportions, particularly the well-formed lunate surface with a deep and narrow acetabular notch, and the complete absence of pachyostosis and osteosclerosis, mark it as a probable middle Eocene protocetid cetacean. Its size corresponds to the newly described Togocetus traversei from the Lutetian deposits of Togo. However, no innominate is known for the Togolese protocetid, which precludes any direct comparison between the two West African sites. The Senegalese innominate documents a new early occurrence of this marine group in West Africa and supports an early dispersal of these aquatic mammals by the middle Eocene.
  

Rodents first appear in the latest Paleocene or earliest Eocene as very fragmentary specimens (Family Paramyidae) known largely from a single locality. After this sparse beginning, rodents are usually abundant in the North American record if proper recovery methods are used. Utilization of rodents for biostratigraphic purposes depends on 1/ extinction, and 2/ replacement by evolution of endemic groups and/or incursions of Old World rodents, and rarely and late by South American kinds. These incursions are separated by relatively long periods of isolation in the Paleogene, but more episodic in the Neogene. At least 10 rodent zones can be characterized by major distinctions, and these zones can be amplified into as many as 16 with little trouble. In general, rodent genera permit as refined a zonation as do genera of large mammals. Distinction at a specific level has not been attempted herein except in the Blancan and Post-Blancan. 

The dentition of Welcommia bodeuri nov. gen. nov. sp. from the Valanginian of Southem France is described and reconstructed. Species and genera of Upper Jurassic and Lower Cretaceous Hexanchiformes are reviewed and discussed.
The genus Notidanoides MAISEY, 1986 must be restricted to the single Nusplingen Upper Jurassic specimen, whose attribution to the species muensteri AGASSIZ, 1843 remains doubtful.
The genus Paranotidanus WARD &THIES, 1987 that does not rest on any type-species nor on any precise dental characterization must be rejected.
The genus Eonotidanus PFEIL, 1983, based on a very poorly preserved and heterogeneous type-material must also be rejected.
Teeth from the Lower Cretaceous, with a peculiar morphology, previously assigned to Eonotidanus or to Notidanoides, are to be ranked in the new genus Pachyhexanchus. 

The hipparions from the Dytiko localities of the lower Axios valley (Macedonia, Greece) are studied. The material comes from three localities Dytiko-l, 2, 3 (DTK, DIT, DKO), which are situated near the village of Dytiko, about 60 km northwest to Thessaloniki. Three species have been determined, the medium-sized H. mediterraneum, the small-sized H. matthewi and the very small-sized H. periafricanum. The determined Hipparion species, their morphological characters and their comparison with the other Axios valley material indicate a Late Turolian age for the Dytiko localities. 

A new species, Otonycteris rummeli nov. sp., is described from the Late Miocene site Gritsev (MN 9) in the Ukraine. Otonycteris rummeli nov. sp. differs from those of most vespertilionids, except recent Otonycteris, Antrozous and Early Miocene Karstala silva, in having a well-developed entocingulid at the foot of the trigonid valley in the lower molars. The morphological resemblance of Otonycteris, Antrozous and Karstala is apparently a case of convergence in the evolution of the Old and New Worlds bat faunas. From at least the Middle Miocene the range of Otonycteris distribution spread to the whole of Central Europe and such a situation continued during the whole Late Miocene. This indicates a more arid climate in Europe during the Upper Miocene compared to the Quaternary. The reduction of the distribution range of Otonycteris and its extinction in most of the territory of Europe could have been caused by the global climatic cooling and increasing glacial cycle amplitude during the onset of the Quaternary. 

Palaeotis weigelti, from the Middle Eocene of central Europe, is a flightless, paleognathous bird. It appears to be a member of the ostrich lineage on the basis of trivial derived characters. It is a very primitive ratite, however, and does not possess any of the highly specialized cursorial adaptations that characterize the modern steppe -and savanna- dwelling ostriches. 

The Paleogene faunal assemblage from Antofagasta de la Sierra (Catamarca, Argentina), is here presented, both in its geological and systematic aspects. The fossil bearing levels are referred to the Geste Formation (Pastos Grandes "Group"). The described specimens belong to the Classes Reptilia (Orders Crocodylia, Serpentes and Chelonii) and Mammalia (three taxa from the Superorder Marsupialia, representatives of the Orders Edentata, Condylarthra, Pyrotheria and Astrapotheria, and six families of the Order Notoungulata). This fauna is referred to the Mustersan Age, which in Patagonia represents the Middle Eocene. Such chronologic assignment is based on the presence of characteristic taxa, their evolutionary stage and on stratigraphic evidence. Finally, a brief comparison with other faunal assemblages from the Early Tertiary of Argentina and Chile, is presented. 

Abstract: A very tiny cuckoo, Chambicuculus pusillus, was recently described after a few fragments of tarsometatarsi from the late Early/early Middle Eocene of Djebel Chambi, Tunisia. New remains, notably a coracoid, confirm the attribution of this genus to the recent family Cuculidae. This coracoid shows a very elongate and strap-like processus procoracoideus. This morphological feature is otherwise only known in the family Cuculidae. The characteristics of the coracoid and tarsometatarsi show that Chambicuculus is morphologically more advanced over the other stem cuculids described in Europe and North America. Chambicuculus is the oldest Cuculidae known so far. 

The first lists of Insectivores (Erinaceidae, Talpidae and Soricidae) from the Pliocene beds of Southern France and North-East Spain are given in this paper. The material from twelve localities is studied. These localities are geographically situated in Languedoc (Celleneuve, Vendargues, Nîmes, Sète, Balaruc 2 and Seynes), in Roussillon (Terrats, Serrats-d'en-Vacquer, Château d'eau and Mont-Hélène) and in North-East Spain (Layna, Medas Islands and Puebla de Valverde). These faunas correspond to the Early, Middle and Late Pliocene. 1 to 8 taxa are identified in these localities and 14 specific taxa are presently listed for this period in this area. Two new specific taxa are described as Galerix depereti nov. sp. from all the Early Pliocene localities in the North-Pyrenean area and as Desmanella gardiolensis nov. sp. from Balaruc 2. For this small mammals, two faunal assemblages are recognized. The first one is dated from the Early Pliocene (F 1, 2 and 3 zones in Aguilar et Michaux) and is characterized by Galerix depereti and rare and little diversified Soricids. The second one is Late Pliocene in age (zones G 2 and G 3). The fossils of the genus Talpa are relatively abundant and the Soricids are diversified and very abundant. The Middle Pliocene (zone G 1) is a transitional period. ln these faunas, most of the insectivore genera are known from the European Late Miocene beds (8 on 10). This fact demonstrates a relative continuity between the invectivore faunas from the Late Miocene to the Early Pliocene. In conclusion, somme paleoecological considerations are suggested.
  

The knowledge of the Anchilophini has been lately renewed by the discovery of a rather large amount of new material still largely unpublished. This new material offers the opportunity of a systematic revision of this tribe gathering those of European Eocene Equoidea which bear no mesostyle on upper check teeth and display a heavy trend to the molarization of premolars.

A cladistic analysis has made out two genera, Anchilophus (Paranchilophus included as a subgenus), characterized by a marked lophodonty and the transverse narrowness of the cheek teeth, a rather high hypsodonty, the frequent occurrence of "crochets" and "anticrochets" on the superior ones, and a rather weak molarization of the premolars, opposite to Metanchilophus n. gen. whose cheek teeth are more transversally elongated, less high, less lophodont, with cusps better distinct, enamel thicker and premolars more molarized on the whole.

Three species of Anchilophus are recognized, A. desmaresti, type species of the genus, A. (Paranchilophus) remyi and A. (Paranchilophus) jeanteli n. sp.

The genus Metanchilophus is more diversified with the species dumasi, radegondensis, gaudini (whose a new sub-species M. g. fontensis is defined), depereli, castrensis n. sp. and chaubeti n. sp.

The skull anatomy has been moreover described with several taxa; it brings to light (for all that one can generalize) that Anchilophini were light animals with a slender and elongated snout, a thin zygomatic arch, a rather developed encephalon with an advanced gyrencephaly.

The structure of the nasal opening together with the occurrence of epitympanic sinuses and the molarizing process of the premolars corroborate the attribution of this tribe to the family PalaeotheIiidae. 

Revision of the Tayassuidae from the old collections of the Quercy leads to interesting facts about taxonomic and phylogenetic problems. It could not have been well done without constant comparison with contemporaneous Tayassuidae from stratified series, as well as with European Miocene Tayassuidae and Hyotheriinae. The result is that only three species are present in the Phosphorites of Quercy: Doliochoerus quercyi FiLHOL (represented by two sub-species: the older, D. quercyi quercyi and the more recent, D. quercyi aquensis), Palaeochoerus gergorianus CROIZET et BLAINVILLE and Palaeochoerus purillus nov. sp. The two former species disappeared with Stampian time, while the third is the direct ancestor of the Aquitanian Palaeochoerus typus POMEL, which became extinct before the beginning of Burdigalian.

The comparison with Hyotheriinae brought to light new criteria for determinations, based on the study of lower premolars. These observations are condensed in a table of keys for determinations.

As a result of these studies, it came to light that the Suoidea of the Oligo-Miocene of Europe have never given rise to, except in rare cases, very long lineages. The different forms of pigs are replaced constantly in Europe by the simple game of extinctions and migrations. The reason of that discontinuity is considered here. 

The litoptem Proterotheriidae are extinct endemic South American ungulates frequently used as an example of evolutionary convergence with the horses. They were considered to be exclusively Tertiary representatives with the youngest record being in the late Pliocene, before the appearence of the equids and cervids during the Great American Interchange. Two undoubted Pleistocene records in Argentina and the specimen here described from Uruguay, confirm the persistence of the proterotherids into that period. In the Quaternary, these ungulates are found outside the typical pampean region and probably were confined to a few northern and warmer more forested relictual microhabitats.

  

Revision of the hipparion material from Montredon, including newly excavated and other unpublished specimens brings evidence of specific heterogeneity.
A fragmentary very small MC III seems very close to H. macedonicum from the upper Vallesian of Ravin de la Pluie, Greece. In that same site was also found a large hipparion.
Most of the Montredon material is referred to H. depereti. This species associates characters usually found in Vallesian hipparions (highly plicated upper cheek teeth, deep vestíbular grooves on the lower cheek teeth, robust metapodials) with characters more frequent in Turolian forms (middle size, lack of confluence in the upper premolar fossettes, lack of ectostylids on the adult lower cheek teeth, well developped keel on the MC III, facette for the 2nd cuneiform present on all MT III). H. depereti shares some of these characters with the Spanish and Portuguese hipparions transítional between the Vallesian and the Turolian (Masia del Barbo, Azambujeira) but is not identícal to any of them. The upper Vallesian hipparion material from Diavata, Greece, probably belongs to H. depereti but not the large hipparion rests from Ravin de la Pluie.
Thus, Montredon and Ravin de la Pluie may well share the same small species, H. macedonicum, but they differ in the associated one: middle-sized H. depereti at Montredon, large-sized H. primigenium at Ravin de la Pluie. Both sites, however, give evidence of small hipparions during the Vallesian, coexisting with other larger species.
The present paper also proposes an adaptation of the Kiesewalter's indices that calculates the height at the withers using the metapodial length, and discusses indices proposed by Gromova and by Sen et al. to express the relative development of the metapodial keel. 

In the course of the current revision of adapid primates from the Eocene Geiseltal, an almost complete specimen was found in the Geiseltal Museum collections. The fossil, the most complete adapid specimen so far discovered in Europe, has been determined as Pronycticebus neglectus n. sp.
Ecology and locomotion as well as the likely phylogenetic position within the infraorder Adapiformes are discussed. 

La faune de Rongeurs d'Escamps (Lot) bien que relativement pauvre en espèces (dix) s'avère riche d'enseignement pour les Rongeurs de l'Oligocène inférieur d'Europe Occidentale. Cette periode semble caracterisée par une cladogenèse des Théridomyines. A Escamps, un nouveau genre (Patriotheridomys) est décrit ainsi qu'une nouvelle espèce de Theridomys. Avec Oltmamys platyceps, décrit ici plus complètement et désormais bien situé stratigraphiquement, les deux espèces précédentes constituent un ensemble original du Sud de la France. A la même époque, en Angleterre, Allemagne et Espagne, des espèces «régionales» de Théridomyinés se dersifient.  A côté de ces formes qui ne semblent pas franchir la «Grande Coupure» sont représentées deux lignées d'Issiodoromyinés (Elfomys sp et Pseudoltmomys cuvieri), une de Théridomyiné (T. (Blainvillimys) rotundidens) ainsi qu'un Gliridé, Gliravus priscus (que l'on différencie nettement de la deuxième lignée des Gliravus oligocènes : G.  meridionalis -> G. majori). On retrouve ces formes dans les gisements plus récents du niveau de Hoogbutsel où elles sont à peine plus évoluées. Le fait que les degrés évolutifs de ces lignées soient très proches laisse supposer que le laps de temps séparant les niveaux d'Ecamps et Hoogbutsel fut relativement court. 

Valverde's cenogram method is used to analyse mammalian communities from the late Eocene to late Oligocene of southern France, mainly from the "Phosphorites du Quercy". Cenogram analysis involves plotting the size of each component species in a fauna on a semilog diagram in rank order, permitting fossil faunas to be compared with Recent ones. The configurations of Recent communities serve as models for establishing the general environmental characteristics of fossil mammalian faunas. This method of analysis applied to faunal sequence can reveal major and sudden ecological perturbations. The paleobiogeographical event (i.e. the mammalian immigration wave) at the Eocene-Oligocene Boundary in western Europe, known as the «Grande Coupure", is here shown to represent a drastic and sudden ecological change: late Eocene tropical environments in Europe deteriorated rapidly turning to subdesert or desert environments al the beginning of the Oligoccne. 

The discovery of new hippopotamid material from the late Miocene Baynunah Formation (Abu Dhabi, United Arab Emirates) has prompted the revision of the existing material of this as yet unnamed fossil taxon. The Baynunah hippopotamid appears to be distinct from all other contemporary and later species in having a relatively more elongate symphysis, a feature similar to the earlier (and more primitive) Kenyapotamus. Yet, the Baynunah hippopotamid presents a dentition typical of the Hippopotaminae. It is therefore a distinct species attributed to the later subfamily, described and named in this contribution. This species provides further evidence for a ca. 8 Ma evolutionary event (termed “Hippopotamine Event”) that initiated the spread and ecological significance of the Hippopotaminae into wet habitats across Africa and Eurasia. The morphological affinities of the new species from Abu Dhabi suggest that the Arabian Peninsula was not a dispersal route from Africa toward southern Asia for the Hippopotamidae at ca. 7.5 Ma to 6.5 Ma. 

A systematic analysis of an hipparionine horse assemblage from Maragheh, Iran is made. A brief orientation to systematic philosophy and informal superspecific characterizations of some Old World hipparionines is given as a background to this work. A character state analysis of skulls is made, and has revealed five distinct species. A character state and stratigraphic trend analysis of isolated check tooth and postcranial remains, with known provenance, is also made. These two combined analyses reveal that the most resolute discrimination of hipparionine species and their evolutionary relationships occurs when multiple character complexes of associated skulls, maxillary and mandibular dentitions are made. When this is not possible, skulls have provided the best basis for discriminating species and their evolutionary relationships. Traditional characters of isolated cheek teeth and postcranial remains are shown here to offer limited information content for hipparionine phylogenetic systematics. The systematic portion of this study includes a comprehensive description of cranial and postcranial remains, and has further corroborated the distinction of five species which belong to at least three superspecific groups including: «Hipparion» geltyi sp. nov., Group 1; Hipparion prostylum (s. l.), and Hipparion campbelli sp. nov., Group 3; «Hipparíon» aff. moldavicum and «Hipparion» ?matthewi, Group 2. These species stratigraphic ranges and evolutionary relationships are also given here and argued to be important for establishing future hipparionine geochronologic correlations between a number of Eurasian late Miocene provinces.