This paper describes the first hexanchid tooth from the Tithonian (Late Jurassic) of New Zealand. For the moment, this tooth represents the earliest representative of the fossil genus Notidanodon in the world and one of the most ancient neoselachians in the Southern Hemisphere. Despite the perfect state of preservation of the unique tooth, the species is left in open nomenclature, pending the discovery of additional specimens. Few nominal species have been assigned to the genus Notidanodon. Four from Cretaceous deposits: N. antarcti Grande & Chatterjee, 1987, Notidanodon dentatus (Woodward, 1886), Notidanodon lanceolatus (Woodward, 1886), Notidanodon pectinatus (Agassiz, 1843) and only two from Paleocene: Notidanodon brotzeni Siverson, 1995, and Notidanodon loozi (Vincent, 1876). Considering the important morphological variations observed between some of these species, it seems obvious that the genus Notidanodon is not monophyletic and will need a revision in the future. 
  

A recently discovered, almost complete specimen of a hybodont tooth, allows us to describe as a new species, fairly common, but usually fragmentary teeth in the Navesink Formation of New Jersey: Hybodus novojerseyensis nov. sp.
Three additional taxa are also described : Heterodontus creamridgensis, Squalicorax sp. and Pseudocorax affinis which is noted for the first time in North America.
The whole elasmobranch fauna of the Early Maastrichtian localities of New Jersey is reevaluated and several generic assignments are changed in comparison to the list previously published in 1975 (Cappetta
& Case). 

Stratigraphic studies in the Aktau Mountains bordering the Dzhungarian Alatau Range in southeastern Kazakhstan have included mapping of Tertiary lithostratigraphic units, documentation of fossiliferous deposits, correlation of sections, etc. These investigations have led in turn to revised interpretation of the Tertiary geology of the area. The Tertiary sequence in the Aktau Mountains is represented by three lithostratigraphic units (in ascending order): (1) the middle Eocene Akbulak Formation; (2) the Oligocene Aktau Formation with a lower member including white quartz sands that contain fossil mammals, and an upper member including red-colored clays and sandstones, brick red clays, an anhydrite and gypsum clayey horizon, and bright brown-red clays; and (3) the upper Oligocene-Miocene Chul'adyr Formation with a lower member of greenish and yellowish conglomerates and gritstones, a middle member including grayish and yellowish sands and gritstones, and an upper member including brown and red clays and carbonate- and anhydrite-rich clays. The Aktau and Chul”adyr Formations represent separate cycles of sedimentation. Mammalian biostratigraphy and biochronology of the three vertebrate faunas in Aktau Mountains are reviewed. The mammalian fauna from white sands of the lower Aktau Formation is small but includes Ardynia and is thought to be early Oligocene in age. The mammalian fauna from conglomerates and gritstones of the lower member of the Chul”adyr Formation is also small but includes Paraceratherium and is thought to be late Oligocene in age. The mammalian fauna from sands of the middle member of the Chul'adyr Formation is extensive, with micro- and macrofauna attributed to Neogene mammal zones MN4 to MN 6, indicating a latest early Miocene to earliest middle Miocene age (Orleanian-Astaracian). Most genera of middle Chul”adyr mammals are known from the middle Miocene Shanwang faunas of China and from the Castelnau-d”Arbieu faunal assemblage (MN4-MN6) of southwestern France. 

Investigations of the Clinchfield Formation (middle Eocene, upper Bartonian) exposed at the Hardie Mine (Wilkinson County, Georgia, USA), produced 4,768 actinopterygian otoliths representing 14 taxa and increased the number of bony fishes threefold from the site. The somewhat limited richness was characterized by bonefishes, mud eels, conger eels, sea catfishes, cusk-eels, snooks, grunts, drums and croakers, and porgies. The assemblage had a relatively even distribution with Ophidiidae, Congridae, and Sciaenidae most common. Included in the otolith taxa was a new sciaenid genus and species, Eosciaena ebersolei, with unknown relationships to other Sciaenidae. The Clinchfield otoliths were compared to other middle and late Eocene in age otolith assemblages in Alabama, Mississippi, and Louisiana utilizing percentage similarity measurements. Analysis indicated that the Clinchfield otoliths were not greatly similar or greatly unlike the Moodys Branch and Yazoo Clay otolith assemblages. However, the Clinchfield showed little relationship to the slightly older Lisbon Formation in adjacent Alabama and is postulated to be related to global climatic and plate tectonic events. Biostratigraphically, the Clinchfield otolith taxa are essentially the same as the other formations except for the Lisbon, which has at least ten unique species. Abundances of Clinchfield otolith taxa indicate a possible sub-bioprovince in the eastern Gulf Coastal Plain. The Clinchfield otoliths indicate a tropical to perhaps subtropical, soft substrate, mainly normal marine to slightly reduced salinities, inner shelf (0–20 m) paleoenvironment with indications of proximal continental coastlines. This investigation represents an initial step in addressing the immensely understudied Paleogene otolith assemblages in Georgia.
  

Welcome to the 18th conference of the EAVP, the first online meeting of our association. The pandemic emergency made it impossible to organize the in-person meeting in Benevento as we all had hoped. However, we couldn’t miss another EAVP meeting. Therefore, this year we are meeting online, trying to make the experience the closest to the in-person meeting possible, in order to offer the delegates the opportunity to share knowledge, build new networks and reinforce the old ones. We have received 137 communications, with more than 150 delegates from 24 countries. All the abstracts have passed a peer review process and are part of this special volume of Palaeovertebrata, the official journal of the EAVP. This year we are also offering a variety of workshops, roundtables and symposia on different topics. These include the annual “Pride EAVP: An LGBTQ+ Roundtable” and “Women in Palaeontology Roundtable Discussion”, together with the workshops on “Gendered Perspective in Palaeontological Research: from Definition to Action”, “International Palaeontology Education: Virtual Teaching and Real-World Learning”, “Stepping out of Academia: Why, When and How?”, “Introduction to Hypothesis Testing in Statistics”, “The Early-Middle Pleistocene Transition: Marked Mammal Turnover and Ecosystem Dynamic” (included in the early event for the XXI INQUA Congress in Rome 2023, “A Mediterranean Perspective on Quaternary Sciences”). To conclude, we are hosting two symposia on “Palaeoart: Diversity on and behind the Canvas” and “3D fossils, Robotic and Experimental Palaeontology”. We wish you all a happy and productive meeting. And see you in Benevento next year! 

The middle Palaeocene of São José de Itaboraí (State of Rio de Janeiro, Brazil) has yielded a very rich and diverse snake fauna which includes Madtsoiidae, Aniliidae s.l., Boidae, Tropidophiidae s.l., Booidea incertae sedis, and Russellophiidae. The present article (part I) deals with Madtsoiidae and Aniliidae s.l. Madtsoiidae are represented by many vertebrae and a few skull bones. They comprise one new species assigned to the genus Madtsoia (M. camposi sp. nov.). However, the definition of the genus Madtsoia is unsatisfactory and the generic allocation might be provisional. A few elements, vertebrae only, belong to the Aniliidae s. l. Two taxa are referred to this latter group: Coniophis cf. C. precedens and Hoffstetterella brasiliensis gen. et sp. nov. The forthcoming part II will deal with Boidae.

  

Pangolins have never shown a high taxic diversity and their fossil record is scarce. We report here the first discovery of a partial humerus from late Oligocene deposits in Pech du Fraysse (MP28, France). The new specimen from Pech du Fraysse is described and compared to various extant and extinct species of pangolins. It shows a suite of morphological features very similar to the humeri discovered in Saint-André (MP 26), Solnhofen (Burdigalien), and Saulcet (Aquitanian), attributed here to Necromanis franconica. The description of the specimen from Pech du Fraysse allowed us to discuss the systematics of Paleogene and Neogene pholidotans. The differences between PFY 4051 and N. franconica on the one side, and N. quercyi on the other side, might be sufficiently important to justify a generic distinction. A comparison with extant species showed that N. franconica was likely terrestrial and fossorial based on its humeral morphology. 

The revision of the Rhombodontidae from the Maastrichtian of Morocco led us to the description of a new species: Rhombodus andriesi.

The biometrical study of populations of R. binkhorsti and R. microdon, species sometimes considered as synonymous, supports the conclusion that they represent two distinct species and not young and adult specimens of a single species. The stratigraphical range of these two species confirms this
result.

The reexarnination of type-series studied by C. Ararnbourg led us to revise the generic status of Rhombodus bondoni which is now ranged, because his dental features, in the new genus Dasyrhombodus showing a less derived dentition than Rhombodus. The stratigraphical range of each species is clarified within the phosphatic series. 

Lalieudorhynchus gandi gen. nov. and sp. nov. is a new caseid synapsid from the Permian of the Lodève Basin, Occitanie, France. This new taxon is represented by a partial but well-preserved postcranial skeleton, and is characterized by the following apomorphies: a transverse section of the sacral and anterior caudal neural spines with a very thin keel-like process anteriorly, a slender dorsal tip of the dorsal and caudal spines, a narrow distal end of the first sacral rib, a fossa on triceps process of metacoracoid, and a very large distal tarsal 1 of same width than the astragalus, with nearly all sides being shallowly concave.
The skeleton corresponds to a sub-adult individual that was excavated from the La Lieude Formation dated as Roadian-Capitanian (Guadalupian). A sedimentological and taphonomical analysis of the type locality, together with preliminary osteohistological observations, suggest that this new French caseid was rather aquatic, as already hypothesised for other large forms.
A phylogenetic analysis of caseids is performed to test the position of this new taxon and to better understand the evolution of the clade: interestingly, Lalieudorhynchus gandi gen. nov. et sp. nov. is closer to the NorthAmerican “Cotylorhynchus” hancocki than to the other French caseids Ruthenosaurus and Euromycter from the Artinskian of the geographically closer Rodez Basin. These two last caseids document the Artinskian radiation of the clade, which remained diverse until Olson’s extinction. Caseids survived, as Lalieudorhynchus is one of the youngest representatives of the clade, and may have used novel ecological strategies to access their vegetarian food sources.

  

Since the 19th century, thousands of footprints were observed in the geological series of the French Mesozoic. All are located in the Triassic and Jurassic. After a promising beginning, in France, it is only a few papers which will be published in the first half 20th century, unlike the USA and of others countries of Western Europe. One ought to wait about 1950 for a revival and now they are nearly 200 papers which were devoted to the ichnofossils. The literature abundance and the renewed interest of the naturalists for the palichnologic studies decided to us to write a synthesis work. This one begins with a stratigraphic inventory in which, localisation, age and paleontological contents of about 180 fossiliferous sites are specified. After having pointed out the followed methods, the footprints paleontological interpretation is then approached in detail and the results obtained are replaced in stratigraphy to deduce the fauna evolution during the Mesozoic. So, it appears that Ichnologic data, more varied and rich in the Triassic and Liassic than those relating to the bones, very rare for the considered periods, are very informative. The middle Triassic (Anisian-Ladinian), thus reveals Cotylosauria, Lepidosauria, Crurotarsi with Rauisuchia, Ornithosuchidae, Crocodylia and Dinosauromorpha more the "Prodinosauria": Dinosamiforme whose skeletons are known in Argentina but only in Ladinian. The rather fast domination of Dinosaurs during Norian is also as well shown. The almost exclusive presence of their footprints, up to fifty cm long, in the Lower Hettangian indicates their supremacy in the environments. Footprints characterise not very deep life places located between inter-supratidal limits and often out of water. Sedimentologic and Palaeontologic studies showed that they were great coastal spaces during Middle Triassic, flood-plain with sebkhas while Upper Triassic, and a large !!coastal marsh!! in Grands-Causses during Liassic in which, mainly, fine stromatolithic layers were deposited. During the same periad, bay beaches spread in Vendée. During the Middle Jurassic, they are also brackish to lacustrine environments and recifallagoons in- the Upper Jurassic. Numerous measurements of the footprints and trackways directions showed that the animaIs moved there in weil defined directions, for long periods. They seem due to the palaeotopography of the life environments relatively stable. Also, the discovery of vegetal radicular networks and small footprints far away from the continental borderlands has suggested that the animals continuously lived in these palaeoenvironnements, belonging to large ecosystems, where the sedimentation rate was weak. This explains that thebadies could not fossilize there but only their footprints through the cyanobacterian action in main cases. From the vertical distribution of different ichnospecies, defined with adapted statistical methods, explained in this work, a palichnostratigraphy was established for the Middle Triassic. Although the footprints are also abundant in Hettango-Sinemurian of "Grands-Causses" and the Vendée, it was not possible, up to now, to establish any zonation in this series; Probably because the palichnofauna is too little diversified there, currently reduced to a majority of Theropods II-IV tridactyl traces.
  

Micro-CT imaging is an increasingly popular method in paleontology giving access to internal structures with a high resolution and without destroying precious specimens. However, its potential for the study of hypoplasia defects has only recently been investigated. Here, we propose a preliminary study to test whether hypoplastic defects can be detected with micro-CT (μCT) scan and we assess the costs and benefits of using this method instead of naked eye. To do so, we studied 13 fossil rhinocerotid teeth bearing hypoplasia from Béon 1 (late early Miocene, Southwestern France) as positive control and 11 teeth of the amynodontid Cadurcotherium (Oligocene, Phosphorites du Quercy, Southwestern France), for which enamel was partly or totally obscured by cement. We showed that all macroscopically-spotted defects were retrieved on 3D reconstructions and selected virtual slices. We also detected additional defects using μCT scan compared to naked eye identification. The number of defects detected using μCT was greater in the Cadurcotherium dataset (paired-sample Wilcoxon test, p-value = 0.02724) but not for our control sample (paired-sample Wilcoxon test, p-value = 0.1171). Moreover, it allowed for measuring width and depth of the defects on virtual slices (sometimes linked to stress duration and severity, respectively), which we could not do macroscopically. As μCT imaging is both expensive and time consuming while not drastically improving the results, we recommend a moderate and thoughtful use of this method for hypoplasia investigations, restricted for instance to teeth for which enamel surface is obscured (presence of cement, uncomplete preparation, or unerupted germs). 

The Causses" is a near 3400 km2 large plateau located in the south of France. Here the first dinosaur footprints where found in 1935. After this, this area has yielded an ever-increasing number of ichnites now in excess of 500 specimens. These latter, 15 to 50 cm long, tridactyl or tetradactyl footprints of generally biped animals, were discovered at the surface of Hettangian to lower Sinemurian dolomite layers within 4 distinct stratigraphic units. The 35 sites bearing ichnites are located on the plateau margin. For the first time, morphologic characters studied through descriptive statistic methods with the usual parameters and classical Student and Snédecor tests, allowed us, to divide the whole set of biped traces into 6 ichnospecies. Their definitions are further constrained by multivariate statistical results using Principal Component Analysis (PCA), Factor Analysis of correspondances (FAC) and Discriminant Analysis (DA). All have confirmed the morphologic observations. So that now, the following taxa are identified : Grallator variabilis, G. lescurei, G. sauclierensis, G. minusculus, Eubrontes giganteus, Dilophosauripus williamsi, cf. Moraesichnium, Orníthopus fabrei nov ichnosp. The more immediately visible differences relate to the interdigital II-IV divarication and the digit length ratio. To this panel, we must add Batrachopus deweyi and shapes suggesting Trisauropodichnus and/or Anomoepus. Among all ichnite associations described in the lower Liasic, the New England assemblage presents the most affinities with ours. It shows the ichnotaxa Grallator, Dilophosauripus, Eubrontes, Batrachopus without forgetting Ornithopus fabrei nov. ichnosp. which is close to Ornithopus gallinaceus from the Massachusetts and Connecticut basins. On comparing the present early Jurassic ichnofauna of the Causses with the ones of the Middle and Upper Triassic formations of the eastem border of the Massif Central (France), it appears that tridactyl footprints become more and more numerous and large from Triassic to Early Jurassic. In the Causses, these latest are prevalent but in Quercy (France), Poland, Italy, USA, they are also associated with Omithopoda, Thyreophora and Sauropoda ichnites. Footprint areas considered here were generaly under an arid climate. Animals that passed by were heavy and bulky possible Megalosaur trackmakers, and lighter and slender Coelophysids or Ceratosaurs. For all, these areas were pathways as the orientations of the trackways seem point out. The directions followed by these reptiles were without any important variation during the Hettango-Sinemurian stages. These areas were also used from time to time by Crocodilomorpha and may be tetradactyl (I-IV) bipedal avian Theropods. However, the number of such trackways in sites, sometimes substantial, should not lead us to overestimate the trackmakers populations. These last were probably relatively moderately abondant in this inter-supratidal swamp environment. In the Causses, ichnites are connected with former algo-laminated deposits (Algal mats) which were rapidly hardened by means of calcitisation of cyanobacteria. The result has been a moderate depth of footprints; autopodia disturbing only a few cm of the carbonate substrate. Other fossils have been discovered : invertebrates with thin bivalve and gastropod shells, crustaceans tests and plants. These latter suggest the existence of paleomangroves like environments but also continental vegetation periodically overruning the swamp environment during regression/transgression cycles. At these times, wooded parts of it, could become protecting, feeding, resting and nesting places.

  

Based on fossil discoveries and phylogenetic studies, an Eocene Asian origin for hystricognathous rodents and anthropoid primates has gained strong support in recent years. The two groups then invaded both Africa and South America, which promoted their evolutionary success. However, the fossil record has so far failed to constrain the nature and precise timing of these pivotal dispersal events. In Africa, given the apparent absence of hystricognaths and anthropoids in early to early middle Eocene localities, it is suggested that these mammal groups dispersed from Asia to Africa sometime during the middle Eocene. In this paper, we report the discovery of several isolated teeth of a rodent from a new vertebrate locality situated in central Tunisia (Djebel el Kébar, KEB-1), dating from the late middle Eocene (Bartonian, ~39.5 Myr). These fossils document a diminutive new species of Protophiomys (P. tunisiensis nov. sp.), a basal genus of hystricognathous rodents which is well known from several North African mammalian-bearing localities of the end of the Eocene. The teeth of P. tunisiensis display a suite of anatomical details comparable with those observed in the other species of the genus, but with a lesser degree of development. Such an apparent primitive evolutionary stage is corroborated by the greater antiquity of this Tunisian species. P. tunisiensis nov. sp. is so far the most ancient representative of hystricognaths in Africa. However, it can be expected that hystricognaths were already present on that landmass given the new data on early caviomorphs recently reported from South America (at ~41 Myr). The arrival of hystricognaths in Africa from South Asia certainly predates the depositional period of the Kébar sediments, but perhaps not by much time. 

The Oligocene deposits of the Ulantatal area in Inner Mongolia (China) contain among the richest mammalian faunas from Asia. To date, only some parts of the rodent faunas have been described. Here, we propose to review the rodent faunal lists for each site, including the description of a few new rodent specimens. We describe three additional rodent species: the Cylindrodontidae Anomoemys lohiculus, the Eomyidae Asianeomys sp., and the Dipodidae Litodonomys huangheensis. This study allows us to constrain the stratigraphic range of Anomoemys lohiculus, which ranged from the late Early Oligocene to the early Late Oligocene in this area. Asianeomys sp. and Litodonomys huangheensis are dated from the latest Oligocene. These Oligocene deposits consist now of more than 70 species of mammals if we include the fauna from Kekeamu. This latter corresponds to the basal part of the Ulantatal Formation and could be dated biochronologically from the earliest Oligocene. When compared to the faunas from the Valley of Lakes in Central Mongolia, the Ulantatal faunas present a great majority of rodents, and this difference can be partly explained by sampling and description biases regarding macro-mammals. This study also shows that variations existed between Inner and Central Mongolia, especially regarding the composition of the rodent paleocommunities. However, the assessment of their evolutionary history in this part of Asia with respect to the important climate and environment changes, require further precisions and more material than current data allow. 

Rhinoceros Giants: the Paleobiology of Indricotheres. Donald R. Prothero. Life of the Past Collection, Indiana University Press; 160 pp. (66 b&w illustrations). Hardback (7x10”): USD 42.00 plus shipping. ISBN: 978-0-253-00819-0. E-book: USD 34.99. ISBN: 978-0-253-00826-8.
  

An interpretation of the marine reptile fossil record, based on the existing litterature and complemented by the review of ancient collections and the study of new material, permits a better understanding of Mesozoic marine ecosystems. An inventory of the marine reptiles known from the Lower Triassic to the Paleocene is presented: 46 families, about 200 genera and 400 species have been recorded. This data base includes commentaries about systematics, stratigraphical ranges and geographical distribution of taxa. Marine reptiles include a mosaic of not necessarily closely related groups: ichthyosaurs, thalattosaurs, hupehsuchians, pachypleurosaurs, placodonts, nothosaurs, plesiosaurs, pliosaurs but also crocodiles, lizards, snakes, turtles. The diversity studies reveal that the fossil record of marine reptiles has been punctuated by two mass extinctions, during the Middle-Upper Triassic transition and at the Cretaceous-Tertiary boundary. The Jurassic-Cretaceous and Cenomanian-Turonian boundaries (the latter marked by the disappearance of ichthyosaurs) are potential crisis periods, but current data are not sufficient to reach conclusions. The Ladinian-Carnian transition is characterized by the disappearance of 64 % of families and affects essentially coastal forms. This extinction coincides with an important regressive phase. During the Upper Triassic, a faunal reorganisation within marine reptiles leads to the progressive disappearance of near-shore forms and to the development of pelagic groups. During the Maastrichtian-Danian crisis, 36% of families died out. Large-sized pelagic forms such as mosasaurs and elasmosaurs were the most affected and their extinction seems to have been rather sudden. On the other hand, pliosaurs and protostegid turtles became extinct, but were already declining. The survivors were near-shore forms such as crocodiles, snakes and some turtles and they may have taken refuge in freshwater environments. A break in the food chain based on phytoplankton is proposed as an extinction scenario for pelagic forms.

  

This paper presents the description and the interpretation of recently discovered traces on a Lower Hettangian dolomitic outcrop in the Bédarieux area, Southern France. One trace set immediately attracted the attention by its resemblance to a small sauropod pes-manus couple but no trackway was visible. As the other traces have a variety of shapes with no obvious significance, it took a thorough examination of the 3D and sedimentological data to come to the conclusion that most traces likely were sauropod tracks made under diverse conditions. Sedimentological and ichnological data indicate that the tracks have been made in the intertidal zone of a carbonated tidal flat shortly before an emersion period. It appears that that the variety of trace shapes is due to a variety of water depths: the sauropods were punting when the water level was high. The lack of trackways seems due to the combination of an underprint situation, buoyancy effects and the small size of the track-bearing slab. Several hypotheses can be considered for explaining the very small size of the tracks, such as insular dwarfism or the immaturity of the trackmakers. 

Two somewhat “odd” taxa of theridomyid rodents, one formerly known (Bernardia marandati Vianey-Liaud, 1991) and the other new (Idicia vidalenci gen. et sp. nov.) are discussed from a taxonomical and taphonomical perspectives. These two rodents were found at Lavergne, a late middle Eocene (MP16) “phosphatière” from the Quercy (Southwest France). The genus Bernardia, being preoccupied by a scale insect (Bernardia Ashmead, 1881), is here renamed Burgia. We benefit from this nomenclatural change to describe additional new dental specimens of this patriotheridomyine species, including a previously undescribed locus (P4). The other theridomyid from Lavergne, Idicia vidalenci gen. et sp. nov., so far documented by a mandible preserving two teeth (m2-m3) is a new taxon of peculiar occlusal morphology, and whose subfamilial affinities remain unknown. These two peculiar theridomyids recorded at Lavergne are found nowhere else, whether in coeval localities in Quercy or elsewhere in Western Europe. We discuss the possible causes of their unique presence at Lavergne.
  

The late Cretaceous Auca Mahuevo nesting site (Neuquén Province, Argentina) has produced a large number of sauropod eggs, many of them containing the remains of embryos. Research at this site has generated important information about the development of the embryos, the morphology and eggshell microstructure of the eggs, and the reproductive behavior of sauropod dinosaurs. Cranial features present in the embryos have allowed their identification as those of titanosaurian sauropods. Differences in the texture of the sediments that contain some of the egg-clutches have illuminated their nest architecture. Microstructural studies of eggshells have expanded our knowledge of their variability and the incidence of pathologies within a reproductive titanosaurian population. Maps showing the spatial distribution of eggs and clutches. the stratigraphic distribution of the egg-beds, and the sedimentological context in which they are contained, have provided the basis for several inferences about the nesting behavior of these dinosaurs. 

A brief historical account of fossil vertebrate discoveries in the Eastern part of the Paris Basin between the beginning of the nineteenth century and 1950 is given. Other localities discovered since then are presented. A reconstruction of past landscapes is briefly elaborated. A biozonation based on mammals is proposed, from the Late Thanetian to the Middle Bartonian. Paleogeographical considerations are added. Suggestions regarding the search for new marnmal localities are made.