Bats are among the most numerous and widespread mammals today, but their fossil record is comparatively meagre and their early evolution poorly understood. Here we describe a new fossil bat from dental remains recovered from late Early Eocene sediments at Prémontré, northern France. This 50 million-year-old bat exhibits a mosaic of plesiomorphic and apomorphic dental features, including the presence of three lower premolars, a single-rooted p3, short p4 with metaconid, myotodont lower molars and a tall coronoid process of the dentary. This combination of features suggests it is an early member of Vespertilionidae, today’s most speciose and geographically widespread bat family. The Prémontré bat has bearing on hypotheses about the origins of vesper or evening bats (Family Vespertilionidae), as well as crown-group chiropterans.


  

The middle Palaeocene of São José de ltaboraí (State of Rio de Janeiro, Brazil) has produced a rich and diverse fauna of boid snakes. It comprises six or seven species: Hechtophis austrinus gen. et sp. nov., Corallus priscus sp. nov., Waincophís pressulus sp. nov., Waincophis cameratus sp. nov.,"Boinae A", and "Boinae B". Moreover, two dentaries might pertain to either H. austrinus or "Boinae B", or even represent a distinct taxon. Hechtophis austrinus is assigned, with reservation, to the Erycinae. All other taxa are referred to the Boinae. The vertebrae of all taxa have paracotylar foramina, which raises the problem of the apomorphic or plesiomorphic nature of this feature. This fauna also raises the question of the presence of extinct erycine boids in South America, but it does not allow this question to be settled. 

The genus Plagiolophus is documented, almost solely in Western Europe, from the middle Eocene up to the mid Oligocene (MP 12 to MP 25), i.e. more than for 15 MY. Seventeen species are now recorded whose two of them are new, P. ringeadei nov. sp. and P. mamertensis nov. sp. Some anatomical variations and the deflection of certain evolutionary trends justify the distinction of three subgenera, Paloplotherium, Fraasiolophus nov. and Plagiolophus s.s. The genus displays a wide range in size and weight (between 10 and 150 kg). The detailed description of the skull of several species is here given for the first time.
Despite important evolutionary drifts during this long time span, the dentition shows a great structural homogeneity, which renders difficult the determination of fragmentary specimens or isolated teeth. It is characterized by a great heterodonty; premolars are little molarized and present a certain regression through time with paradoxically some progress in the molarization. The hypsodonty increases: the first Plagiolophus are hardly less brachyodont than Propalaeotherium, and the last ones are nearly as hypsodont as Merychippus from the early Miocene. The upper molars change from a wide crown pattern, with an open occlusal surface, lightly oblique transverse lophs and rounded internal cusps, to a narrower pattern, with a frontally constricted occlusal surface and internal lophs aligned parallel to the ectoloph. The M3/3 become always longer.

The dental enamel displays horizontal Schreger-bands with imprecise limits occupying only the middle part of the enamel layer. The dentine is remarkable by its high rate of pericanalicular dentine. The crown cementum, lacking in earlier forms, increases to the point where it fills the occlusal valleys of the
teeth.

The masticatory musculature shows a increasing prominence of the temporal, with probably an important role devoted to the pterygoid muscles in lateral movements related to a two-phase type of chewing.

The evolution of the dentition, of the masticatory musculature and of the repartition of masticatory forces indicate that the Plagiolophus have known different diets through their long evolutionary history; at first browsers they became mixed feeders and finally grazers. Their relatively long neck allowed these animals to reach different vegetal layers. The strength of the nuchal crests also suggests that they were able to have strong backwards movements of the head to pull up their food.

This evolution of diet seems related to the slow degradation of environmental conditions attested during this period in western Europe, with the generalization of more open landscapes, increasing aridity and more marked seasons.

Besides, a remodeling of the face is ontogenetically and along time observed, in relation with the evolution of the masticatory apparatus and especially with that of the mandibular lever arm. The postcanine diastemata become longer in the course of evolution; the free extremities of the nasals are always relatively long which contradicts the hypothesis according to which Paloplotherium may have had a trunk. At last the lineage Fraasiolophus can be distinguished by the presence of a deep malar fossa, probably related to a strong development of the maxillo-labialis superior muscle.

The orbit is always large and tends to increase in size, which indicates a good development of the vision and its increasing role in the life relations. A peculiar type of epitympanic sinus could have been used as a resonance chamber insuring a certain amplification of sounds before their transmission to the eardrum. The endocranial cast reveals a relatively large brain with an advanced degree of gyrencephaly. Beside the role eventually played in food research and social relations, these neurophysiological abilities, also related to an advance in cursorial fitness, could have contributed to the survival of these animals facing the predation pressure of the first fissipede carnivores and the competition with new immigrant herbivores after the "Grande Coupure".
On the basis of some shared apomorphies with the Pachynolophinae, which prevent from considering the latter as Equidae (molarization of the premolars, reduction of the premaxilla dorsal apophysis, peculiar epitympanic sinus, splitting of the jugular process), the hypothesis of an autochthonous origin of Plagiolophus issued from a form near Propalaeotherium, is once again proposed and discussed. Finally, intra-generic relationships are taken into consideration. 

The hipparions from the Dytiko localities of the lower Axios valley (Macedonia, Greece) are studied. The material comes from three localities Dytiko-l, 2, 3 (DTK, DIT, DKO), which are situated near the village of Dytiko, about 60 km northwest to Thessaloniki. Three species have been determined, the medium-sized H. mediterraneum, the small-sized H. matthewi and the very small-sized H. periafricanum. The determined Hipparion species, their morphological characters and their comparison with the other Axios valley material indicate a Late Turolian age for the Dytiko localities. 

Sainte-Néboule has yielded only 4 species of Rodents. But the Theridomyids (Blainvillimys rotundidens and Patriotheridomys altus) are very significative of the age of the locality: Ste-Néboule is lower than the marker level of Escamps 

A strong lophodonty, an extreme heterodonty, some hypsodonty and regular overlayings of coronal cement are prominent features of the genus Leptolophus (Palaeotheriinae = Palaeotheriidae s.s.). The histological pattern of the teeth unusually joins type II enamel prisms, characteristic of advanced ungulates, together with archaic features, such as an almost complete lack of Hunter-Schreger zonation and a weak expanse of peritubular dentine. The skull is narrow and slender, with an elongated ante-orbital facial region, a moderately notched nasal aperture, a rather elongated post-canine diastem, parallel zygornatic arches and a fairly dorsally located squamoso-mandibular joint.The functional analysis brings to light "ectolophodont" masticatory cycles with two phases, in which maximum power was applied, contrary to equíds, on hindmost teeth; likewise, skull accomodations to increasing height of the teeth are quite different. This study leads to the assumption that Leptolophus may have been light mammals, living in rather open surroundings, browsing on herbaceous plants or leaves cropped close to the ground. Moreover, it appears that it could have been some inadequacy of dental structures to the dietary, which leaded to quick wear of the teeth and to many enamel notches, but had been somewhat balanced by the early increase of hypsodonty, not induced in such a case by a biotop deterioration (as it will happen at the end of the Eocene). This ínadaptation might account for the short duration of the genus Leptolophus, whose the 3 species, L. stehlini, L. nouletí and L. magnus n. sp. are indeed confined in the level MP 16. Its geographical spreading (as far as known, South of western Europe) and the morphological pattern of its dentition suggest that this genus would have been related to early upper Eocene endemic spanish forms.





  

The petrosal of the Paulchoffatiidae HAHN, 1969 is described and compared with that of younger multituberculates and of other Mesozoic mammals. The "Morrison petrosal", described by Prothero (1983), is also discussed; it probably belongs to the multituberculates. The reconstruction of the ventral side of the Paulchoffatiinae-skull, given by Hahn in 1987, is completed by addition of the otic and the occipital region. 

The present study of Artiodactyla from the Middle Eocene of the Geiseltal lignite beds concems systematics, biostratigraphy, and palaeoecology on the basis of 174 specimens: isolated remains to more complete skeletons. Instead of the formerly known five species of two families are now recognized 14 species of the Diacodexeidae, Dichobunidae, Cebochoeridae, and Haplobunodontidae. New species are Aumelasia maniai, Anthracobunodon neumarkensis, Masillabune franzeni. Four species of the Geiseltalfauna are definitely known from elswere, and five species are closely related to those from other European localities. Evidently the faunal situation of artiodactyls during the Middle Eocene of Europe was largely uniform. The distribution of artiodactyls within the sequence of the Geiseltal strata corroborates the biostratigraphical concept of the land mammal age Geiseltalian (Franzen & Haubold l986b) as well as the mammalian reference levels MP 11-13 (Franzen 1987). Reconstructions of the skulls and skeletons allow conclusions on the functional morphology and palaeoecology of the artiodactyls of the European Middle Eocene 

The locality of Montredon has provided 8 species of camivores, often little documented, that are discribed and depicted. All the remains are fragmentary and generally badly preserved which lowers very much the possible precision of the taxonomic study; this one has however allowed the creation of a new subspecies. The most richlypdocumented forms are an ursid (Indarctos) and a felid (Machairodus). Issuing often from a westem Europe evolution, the carnivores are well inserted between those of the better known faunas of the "Classical Pontian" and of the lower Vallesian and this situation fits also well with their stratigraphic level, that cannot however be more accurately defined with them alone.
  

Cet article présente une étude détaillée du genre Stehlinia, un chiroptère du Paléogène d'Europe, basé sur un matériel abondant issu notamment du gisement d'Escamps (Quercy). L'analyse révèle que Stehlinia possède un mélange de caractères primitifs (comme une denture tribosphénique complète et des prémaxillaires soudés) et évolués (notamment dans le squelette post-crânien), le rapprochant des vespertilionoïdes actuels, en particulier des Kerivoula. 

A recently discovered, almost complete specimen of a hybodont tooth, allows us to describe as a new species, fairly common, but usually fragmentary teeth in the Navesink Formation of New Jersey: Hybodus novojerseyensis nov. sp.
Three additional taxa are also described : Heterodontus creamridgensis, Squalicorax sp. and Pseudocorax affinis which is noted for the first time in North America.
The whole elasmobranch fauna of the Early Maastrichtian localities of New Jersey is reevaluated and several generic assignments are changed in comparison to the list previously published in 1975 (Cappetta
& Case). 

We report the first confirmed fossil bats from North Dakota, including a new species referable to the Vespertilionidae represented by a maxilla with P4-M3 from the Brule Formation, Fitterer Ranch local fauna, early Oligocene, Whitneyan North American Land Mammal Age. Unassociated postcranial fragments of the humerus and femur also represent a vespertilionoid, but appear to reflect a different, unidentified species. The new taxon, Quinetia frigidaria sp. nov., is referred to the genus Quinetia, previously known only from approximately contemporaneous deposits in Europe. The new species is larger than Quinetia misonnei from the early Oligocene of Belgium. It is similar in some morphological characters to Chadronycteris rabenae (Chiroptera incertae sedis) of the late Eocene (Chadronian) of northwestern Nebraska and to Stehlinia species (?Palaeochiropterygidae) from the Eocene and Oligocene of Europe, but differs from each in morphological details of the dentition and maxilla. An unassociated talonid of a lower molar from Fitterer Ranch shows myotodont morphology, unlike the nyctalodont lower molars in Q. misonnei, and thus represents a second chiropteran taxon in the fauna. Quinetia frigidaria is a member of a Paleogene radiation of bats near the low point of the Eocene-early Oligocene decline in global temperatures, increased seasonal aridity, and loss of tropical floras from mid-latitude North America. 

The Las Águilas site near Porvenir de Jalpa, Coahuila, Mexico, is extremely rich in tetrapod remains comprising both bones and trackways of several dinosaur taxa of late Campanian age. Within a 50 m thick section we identified at least nine layers with dinosaur bone assemblages. In one of these the dinosaur bones are associated with remnants of eusuchian crocodilians, turtles, plesiosaurs, pterosaurs, tyrannosaurids, dromaeosaurids, parksosaurid, hadrosaurids, ceratopsids, and ankylosaurs. This layer is also rich in coprolites of turtles, crocodilians and likely theropods, thus providing evidence for the wealth of Late Cretaceous vertebrate life in the area. 

La faune d'artiodactyles de Ste-Néboule, qui comprend neuf espèces, présente de nombreux
points communs avec les faunes habituellement rattachées au Ludien supérieur, telles celles de La Débruge ou de Montmartre. A l'inverse de ces localités, on ne connaît pourtant, à Ste-Néboule, ni Oxacronae, ni Anoplotheriinae. Parmi les espèces du gisement se trouve une nouvelle espèce du genre Mouillacitherium (M. schlosseri n. sp.), cette forme devant être interprétée comme le dernier représentant du rameau. Nous faisons connaître par ailleurs la presque totalité de la denture du Dacrytherium saturninii, espèce qui était à ce jour très imparfaitement documentée. Ste-Néboule est, d'autre part, la seule localité où l'on peut signaler l'association probable de deux lignées du genre Amphimeryx. Les primates sont peu diversifiés à Ste-Néboule, puisque le groupe est limité au seul genre Adapis. Les genres Microchoerus et Pseudoloris, que l'on sait pourtant être représentés dans des gisements dâge voisin (Microchoerus ornatus à San Cugat de Gavadons et Mormont-Entreroches, Pseudoloris reguanti à San Cugat de Gavadons, Pseudoloris cf. reguanti à Neustadt ; cf. Louis et Sudre 1975) sont absents dans les faunes du Quercy de cette période. 

Of the ten cricetid species from the Oligocene of Western Europe, attributed until now to the genus Eucricetodon, only four prove to be utilizable - E. atavus, E. huberi, E. praecursor, E. collatum - to which it is possible to add two forms newly described: E. huerzeleri and E. quercyi. The evolullon of the genus Pseudocricetodon is also the subject of new observations. The study of the dental morphology allows us to distinguish in these two genera three lineages beginning in the middle Oligocene:

- Lineage P. montalbenensis-P. thaleri (from Montalban to Goderet), of small size, without
increase in size.
- LineageE. atavus-E. infralactorensis (from Hoogbulsel to Estrepouy), of middle size, with a regular increase in size.
- Lineage E. huerzeleri-E. haslachense (level of Montalban to that of Estrepouy), of large size, with an increase in size.

Two other forms are equally represented in these loealitles: Heterocricetodon aff. helbengi and Melissiodon quercyi. It has been possible to attribute a precise age (zone of "Cournon") to the last species, which has been defined by Schaub (l925) from material in the Old Quercy collections. The genera Pseudocricetodon, Eucricetodon, ? "Cricetodon", Leydimys, Eumys, differentiated at the beginning of the middle Oligocene in Europe, Asia and North America, seem to derive from a common ancestral group. The place of origin of this group could be situated in Asia. 

Sieving and washing of about 700 kg of sediment from the miocene site at Montpeyroux produced otoliths of 34 teleost species, of which four still occur in the present day fauna or are near to extant species. Among the fossil species, eight are new : Ilisha lerichei, «genus Clupeidarum ›› orbiculatus, Dipulus mediterraneus, Morone cornuta, Chanda nelsoni, Pomadasys steurbauti, «genus Sciaenidarum ›› barthassadensis and Paraplagusia roseni. The fauna found is typical for a tropical or subtropical very littoral, probably even estuarine environment; it was living in a period near the boundary between Lower and Middle Miocene, perhaps somewhat earlier. 

Designated the type-locality of a late Eocene paleomammal zone, Robiac has recently been the object of important excavations. The first results of the new collecting, as  well as a revision of the material in old collections, are given in this work.
Two stratigraphic section, cutting through the two sites presently distinguished (Robiac-Nord and Robiac-Sud) reveal the lithologic variation at the base of the Fons Limestone and the localization of three fossiliferous beds at Robiac-Sud.
The molluscan fauna and the flora (charophytes), which have already been described, as well as that of the lower vertebrates, have been listed.
A list of 46 mammalian species (only 16 species were known previous to 1964) has been established. The micro-mammals, nearly all new in this fauna (marsupials, insectivores, bats, rodents, primates, and some smallsized artiodactyls), Were obtained only after screen-washing of the matrix; about 4 tons of sediment were thus treated.
The artiodactyls have been the most extensively analyzed; 6 genera, of which one is new, have been recognized. The latter is described as Robiacina minum n.g., n.sp., and represents a very small artiodactyl of the family Anoplotheriidae. The taxonomie status of certain species formerly described has been clarified by the designation of lectotypes (Cebochoerus robiucensis, Catodonerium robiacense, Xiphodon castrense).
The paleontologic corrélations at present possible between the late Eocene faunas have allowed the relative positions of te different French localities of this age to be established; the Guépelle locality, it seems, could define in the future a new paleomammal zone.
  

New dental and skeletal material referable to Palaeochiropteryx tupaiodon from the Middle Eocene locality of
Messel (G.F.R.) is studied, which provides additions to the previously gained knowledge of this european genus. Dental specimens from Geiseltal (G.D.R.), also of Middle Eocene age, allow us to analyze Cecilionycteria prisca. Some of these are separated to establish a new genus, Matthesia, and two new species, M. germanica and M. ? insolita. 

Three new fossiliferous localities, two of karstic origin, Castelnou 3 and Font Estramar, respectively Late Upper Miocene and Lower Pliocene, and one of lacustrine origin, Thuir, Lower Pliocene, add data about the transition between Miocene and Pliocene faunas of rodents in southern France. An unexpected association of taxa was present in the late Upper Miocene, including between others, Myocricetodon, Hispanomys, Ruscinomys, Cricetus barrierei, Promimomys and a new species of Stephanomys, S. dubari nov. sp. Myocricetodon is still known in the Lower Pliocene. It is shown that the large field-mice known since the Late Upper Miocene belong to two different lineages, on one side, A. jeanteti, on the other side, A. gudrunae followed by A. gorafensis. Biochronological and biogeographical implications are discussed. 

The description of new material of the Upper Eoœne of France, Southem Germany and Spain together with the restudy of the material described by Thaler (1966b) as Pairomys allows to confirm the validity of the genus Ectropomys up to know controversally discussed. As a consequence the subfamily Remyinae and Oltinomyinae can be defined precisely. The study of the dental characters shows that the Remyinae in fact belong to the Theridomyidae and represent a derived evolutionary unit within this family. The Oltinomyinae together with the Theridomyinae and Issiodoromyinae very probably form a monophyletic unit.