Messiphoca mauretanica nov. gen., nov. sp. représente le premier Phocidae fossile recensé en Afrique du Nord. Provenant du gisement sahélien (Miocène supérieur) de Raz-el-Aïn (Algérie), il est connu par quelques os du membre antérieur (humérus, radius, ulna), quelques vertèbres dorsales et un crâne très fragmentaire. La description de cette forme nouvelle amène à considérer Messiphoca mauretanica comme un Monachinae archaïque proche de l'origine du groupe Pliophoca - Monachus. L'incidence de cette interprétation d'un Phocidae «pré-Messinien ›› sur la crise de salinité du Messinien est aussi envisagée. 

The fossil mammals discovered in the quarry of Rians (Sparnacian, Provence) are described. Among these forms, Hyracotherium is interesting because of the little molarization of the lower premolars and its small size, and Diacodexis by its small size and very primitive astragalus ; they may be the most primitive representatives of their respective orders. Also, Proviverra eisenmanni n. sp. is the smallest and most primitive hyaenodontid yet described. Hyopsodus itinerans is the first species of this genus described France. Among other rare fossils is a new species of bat, a small palaeoryctid, and other forms not yet identified. Marsupials are varied. Several new species are present among the rodents. The fauna is well-balanced and rich in small hyopsodontid condylarths. It is stratigraphically situated at the
Dormaal reference-level, at the base of the early Eocene, and is considered equivalent to the late Clarkforkian of North America. The hypothesis is presented that new forms appearing at the beginning of the Wasatchian in North America migrated, in fact, at that time from Europe.

  

The biometric and morphologie variability of the cheek teeth in the end-of-the-phylum species Lophiodon lautricense Noulet studied in this note, reposes on the observation of about 800 teeth. These were revealed to be little variable in absolute dimensions. The considerable morphologie variability in the upper premolars permitted the problem of the molarization process to be taken up. An hypothesis concerning the order of eruption of the cheek teeth is formulated based on an examination of a large number of milk dentitions. In conclusion, it is suggested that reservations be held on the value of dental characters classically used in systematics for the group under consideration. 

The new fauna of Collet Redon (Bouches-du-Rhône, France) is dated by three rodents: Megacricetodon aff. bavaricus, Democricetodon affinis mutilus and Peridyromys cf. hamadryas. They correlate this locality with Oggenhof and Ohningen in Bavaria (Western Germany). As the radiometric age of Ohningen is estimated between 14 and 13 M.Y., these three localities are of Serravallian age. This datation brings a complete readjusment of the stratigraphy of the section of Collet Redon formerly described by Collot and Combaluzier. The marine deposits with underly the continental formation with the mammal fauna, are Burdigalian. The angular unconformity between the marine and the continental deposits gives evidence of an episode of emersion on the margin of a sedimentary basin, with deformation and erosion. Owing to the newly discovered fauna, this geodynamical event is clearly settled within the regional geographical and chronological context. Lacustrine and continental deposits of such an age were up to now unsuspected in this area. 

The wear facets of primitive rodents can be homologized with those of primitive primates and ungulates. As in primates, the jaw movement was ectental, with an increased anterior component in the lingual phase (phase ll). The buccal phase (phase I) in rodents approaches the horizontal and it tends to be reduced in importance in comparison with the lingual phase. ln more advanced rodents the efficiency of grinding is increased by the development of additional cutting edges of enamel (e.g. enlargement of hypocone, development of mesoloph and lingual sinus). The buccal phase movement becomes lined up with the lingual phase movement to form a single oblique chewing stroke,resulting in planation of the crown. As the stroke becomes more longitudinal (propalinal) the enamel edges become more transverse. In Muridae propalinal chewing evolved before the loss of cusps, facets were reorientated and additional cusps developed. 

In dilambdodont molars the primitive crest between paracone and metacone (centrocrista) is represented by a pair of crests that join the mesostyle (postparacrista, premetacrista). The cutting action of these crests against the crests of the hypoconid is described. Dilambdodonty is a derived adaptation for greater cutting efficiency. It has evolved several times and in more than one way. 

Many types of insectivores have been described from specimens found in the Quercy phosphorites. These remains very often were not dated because they came from old collections. Recent excavations have permitted the situation of Amphidozotherium cayluxi FILHOL in the late Eocene. Two new genera are descrlbed based on material both from the old collections (Cryplotopos nov. g.) and from that recently recovered (Darbonetus nov. g., beginning of the middle Oligocene). Their systematic positions are revised and comparisons with American faunas are made. Amphidozotherium is not a talpid, but an erinaceoid belonging to an indeterminate family. Saturninia gracilis STHELIN is classified among the Nyctitheriinae, Myxomygale antiqua FILHOL among the Urotrichini Talpinae, and the genus Geotypus POMEL among the Scaptonichini Talpinae. A new study of the talpids from Auvergne has been rendered necessary. During the late Eocene and Oligocene precise morphology relationships existed between certain insectivores of Europe and North America. 

Cet article présente les premières données sur les Carnivores fissipèdes des Phosphorites du Quercy, issues de fouilles récentes (1971-1972). Les fossiles se répartissent de l'Oligocène inférieur à supérieur. 

A systematic analysis of an hipparionine horse assemblage from Maragheh, Iran is made. A brief orientation to systematic philosophy and informal superspecific characterizations of some Old World hipparionines is given as a background to this work. A character state analysis of skulls is made, and has revealed five distinct species. A character state and stratigraphic trend analysis of isolated check tooth and postcranial remains, with known provenance, is also made. These two combined analyses reveal that the most resolute discrimination of hipparionine species and their evolutionary relationships occurs when multiple character complexes of associated skulls, maxillary and mandibular dentitions are made. When this is not possible, skulls have provided the best basis for discriminating species and their evolutionary relationships. Traditional characters of isolated cheek teeth and postcranial remains are shown here to offer limited information content for hipparionine phylogenetic systematics. The systematic portion of this study includes a comprehensive description of cranial and postcranial remains, and has further corroborated the distinction of five species which belong to at least three superspecific groups including: «Hipparion» geltyi sp. nov., Group 1; Hipparion prostylum (s. l.), and Hipparion campbelli sp. nov., Group 3; «Hipparíon» aff. moldavicum and «Hipparion» ?matthewi, Group 2. These species stratigraphic ranges and evolutionary relationships are also given here and argued to be important for establishing future hipparionine geochronologic correlations between a number of Eurasian late Miocene provinces. 

The rodents from the spanish locality of Can Ponsic 1 bring new data about some rodents species of the beginning of the Upper Miocene in South-West Europe. The criticims made by Mein and Freudenthal about the validity of the species Hispanomys thaleri from Can Llobateres are not justiíied. The study of the anatomy of the skull of Rotundomys from Can Ponsic 1 gives accurate information about the affinity of this genus with Cricetulus, and shows that the hypothesis, according to which Rotundomys is an ancestral form of the Arvicolids, is unlikely. The systematics of Heteroxerus and the phylogeny of the mio-pliocene Muscardinus species are also discussed. The Can Ponsic 1 locality is a little older than Can Llobateres.

  

The fluvial sediments of the Siwaliks in northern Pakistan contain a geologic and paleontological record encompassing the Miocene and Pliocene. The sediments, which are thousands of meters thick, have abundant vertebrate and invertebrate fossils from multiple horizons. Fossils were initially discovered in the Siwalik Hills of India in 1832 and there are collections of Siwalik fossils not only in the Museum of the Geological Survey of India, but also in various European, British, and American museums. Renewed collecting in northern Pakistan in collaboration with the Geological Survey of Pakistan has now added over 10,000 bovid fossils to a new collection in Islamabad. The fossils include many isolated horn cores and occasional skulls, as well as teeth and postcranials. Here we discuss material belonging to the Antilopinae, recognizing fifteen species belonging to at least three tribes. Five of the fifteen have previously been named, while an additional two have cranial material suitable for defining a new species. The remaining eight are known from cranial material that allows us to distinguish them from other Siwalik bovids, but in most instances they are of doubtful generic identity and we think may belong to previously recognized taxa from other geographic regions. One (?Protragelaphus sp.) probably belongs to an unnamed species, but because the available material is inadequate for defining a new species, we do not name it at this time. The Antilopini predominate in the fossil collections, both in terms of number of species and number of specimens.
  

The Late Miocene site of Gritsev (MN 9, Ukraine) has yielded a very rich bat fauna, the remains of which are well preserved. Compared to other Neogene bat assemblages of Europe, the Gritsev bat community is unique in preserving plecotine bats, which are rare from Neogene sites. Some peculiar and new bat species, including a large plecotin Otonycteris, already were described from the Gritsev mammal site. Here we report new records of small plecotin bats from Gritsev, including a new taxon, Barbastella maxima nov. sp. This is the earliest reliable fossil record of this genus and it differs from more recent species of Barbastella in being considerably larger. The evolutionary patterns in the odontology within the tribe Plecotini, supported by biostratigraphical distribution of fossil records of Plecotus are discussed. The morphological peculiarities of the new fossils of plecotine bats from Gritsev are discussed in connection with its possible taxonomical affinity. 

Valverde's cenogram method is used to analyse mammalian communities from the late Eocene to late Oligocene of southern France, mainly from the "Phosphorites du Quercy". Cenogram analysis involves plotting the size of each component species in a fauna on a semilog diagram in rank order, permitting fossil faunas to be compared with Recent ones. The configurations of Recent communities serve as models for establishing the general environmental characteristics of fossil mammalian faunas. This method of analysis applied to faunal sequence can reveal major and sudden ecological perturbations. The paleobiogeographical event (i.e. the mammalian immigration wave) at the Eocene-Oligocene Boundary in western Europe, known as the «Grande Coupure", is here shown to represent a drastic and sudden ecological change: late Eocene tropical environments in Europe deteriorated rapidly turning to subdesert or desert environments al the beginning of the Oligoccne. 

The species diversity of the mammalian fauna from Montredon (Hérault, France, late Miocene) is examined in terms of richness and abundance. A cenogramic analysis of the fossil mammalian community suggests the prevalence of open habitats, with the presence of marshes and of a poorly developed galery forest, and a climate rather warm and dry. 

Besides Dendromurids and Murids, the Chorora Rodents include 7 taxa, two of which are new. Afaromys nov. gen. is a Cricetodontine with very complicated lower molars, certainly isolated for a long time from Eurasian species. A Paraphiomys, with tetralophodont upper molars, is also quite different from other East African species, and its roots should probably he searched in the middle Miocene. Other taxa are less original, and more similar to the South Asiatic ones. 

A short review of the genera and species of snakes from the Quercy's phosphorites described by former authors is followed by the study of specimens recently collected. Most of these latter specimens belong to the Aniliidae, Boidae, Colubridae and to the Scolecophidia; the precise systematic position of some of them is not defined yet. The following genera and species are described: Eonilius europae nov. gen. and nov. sp., Platyspondylia lepta nov. gen. and nov. sp., Coluber cadurci nov. sp. and Dunnophis cadurcensis nov.  sp.; a species described by de Rochebrune (Palaeopython filholi) is revised. One of the most important conclusions of this study is that the Colubridae appear as early as the Upper Eocene. The stratigraphic repartition of these snakes shows that the rich Upper Eocene fauna is followed by the very reduced fauna of the Lower Oligocene; then the oligocene «Grande Coupure» corresponds to a very important faunistic renewal. The fauna remains poor and little diversified during the beginning of the Middle Oligocene after which there is a new faunistic explosion. 

Voici une centaine d'années débutait sur les Causses du Quercy l'exploitation des phosphorites qui devait permettre d'amasser des collections de Vertébrés fossiles uniques au monde. Ces fossiles firent l'objet de nombreuses publications et les noms de Filhol, Gaudry, Schlosser, Stehlin, Teilhard de Chardin sont attachés à ces premières études.

Après quelques années de prospection, les fouilles dans le Quercy ont repris en 1965, organisées par les laboratoires montpelliérains et parisiens. Ces premières recherches ont été aidées par le Centre National de la Recherche Scientifique qui a organisé une « table ronde» pour faire le point des premiers résultats obtenus. Ceux-ci ont été jugés suffisamment prometteurs pour que le C.N.R.S, ait décidé de poursuivre son effort par l'intermédiaire d'une Recherche Coopérative sur Programme.



  

The comparative morphology and biometry of the lower cheek teeth of modern Equus are studied on approximately 300 mandibles belonging to the 10 usually recognised species : Equus grevyi, E. burchelli, E. quagga, E. zebra, E. africanus, E. asinus, E. hemionus, E. klang, E. przewalskii, E. caballus. The studied parameters comprise : occlusal length and width, postflexid length and index ; shape of the double knot (metaconid + metastylid + lingual groove) ; depth of the vestibular groove on the molars ; frequency of the pli caballinid, protostylid and other enamel plications or islets ; frequency of the dP/l.
The same methods of study are applied to a number of North American, Eurasian and African species. For the sake of comparison, some Hemphillian equids were observed (Dinohippus interpolatus, Dinohippus leidyanus, Astrohippus ansae, Phiohippus mexicanus) but most of the discussed material belongs to Pliocene or Pleistocene species of Equus : the « stenonine ›› E. stenonis, E. simplicidens, E. sanmeniensis and E. teilhardi; the « caballine ›› E. scotti, E. lambei, E. Iaurentius, E. mosbachensis, E. germanicus, E. gallicus, E. taubachensis and the Liakhov horse. The relationships of other species, in particular the North American E. calobatus, E. occidentalis, E. cf mexicanus are not clear for the moment. ln Africa, the Plio-Pleistocene species from Koobi Fora (Kenya) show some stenonine and perhaps asinine affinities. The relationships of E. numidicus and E. tabeti are uncertain but these species are probably related to the East African ones. E. mauritanicus is most certainly related to the Quagga group.
The biometrical data are gathered in 32 tables ; 4 photographie plates and 19 figures illustrate the next. The whole is a complement of the previously published studies of the skulls, upper cheek teeth, incisors and metapodials of modern and fossil Equus.

  

The Pantolestidae from the middle eocene of Bouxwiller are the subject of a detailed study. Buxolestes hammeli (n. g., n. sp.) is not closely related to any other European or North American form described until now; it presents, however, some characters in common with Pantolestes, a form of the same age from North America. A parallel evolution from a common ancestral form could explain this ressemblance.
Another form (gen. and sp. indet.) accompanies Buxolertes hammeli in the Bouxwiller fauna.
  

The exploitation of new early Eocene localities in the Paris Basin has resulted in the collecting of  numerous mammalian remains, among which are about 300 isolated teeth representing the rodents. They belong, for the most part, to the paramyid group. Only the latest level of the early Eocene has yielded rodents belonging to the pseudosciurid group. The paramyids, the object of this study, are represented by at least 5 genera and 10 species; they are distributed among 4 clearly dilferentiated subfamilies : Paramyinae Simpson 1945, Pseudoparamyinae Michaux 1964, Ailuraviínae n. subf., Microparamyinae Wood1962.
It results from this study that the principal types of rodents in the middle and late Eocene of Europe are clearly related to the forms described here: Plesiarctomys Bravard 1850 is related to Pseudoparamys Michaux 1964, Ailuravus Rütimeyer1891 to Meldimys n. gen., Gliravus hammeli Thaler 1966 to Microparamys nanus (Theilard1927), and some Masillamys Tobien 1954 to Microparamys russelli Michaux 1964 and to M. sp. 1. Gliravus and Masillamys are the oldest representatives of the Gliridae and the Pseudosciurídae, respectively, families which will progressively replace the Paramyidae in the course of the middle and late Eocene.
Two stages can be recognized in the history of te European Paramyidae fauna during the course of the early Eocene: the older is characterized by the Mutigny fauna, the younger by the Cuis fauna.
The Mutigny fauna appears already rather diversified and does not completely correspond, point by point, to the homologous fauna of North America. A geographic differentiation seems to have been manifested rapidly, unless the fauna established in Europe was already a little different from that which established itself in North America.