Encyclopedia of Paleoherpetology. In 19 parts ; Initiated by Prof. Dr. Oskar KUHN ; Edited by Dr. Peter WELLNHOFER, München. - Part. 17 A : Pelycosauria, by Dr. Robert R. REISZ, Erindael Campus, University of Toronto. 1986. VIII, 102 pp., 43 fig., 20,5 x 25,5 cm, soft cover : DM 140, - (Pref. - Price for subscribers to the whole series : DM 126, -). ISBN 3-437-30486-0. 

The genus Cryptomeryx SCHLOSSER, 1886, inusited during a long period, has been discovered in Lower and Middle Oligocene localities of the Quercy region (South-West France). This new material, as well as specimens from the old collections referred to Cryptomeryx, are described; their study, allows us precising the definition of the genus, and confirming its allocation to the Tragulidae family. The type species of the genus, Crypmmeryx gaudryi (= Lophiomeryx gaudryi FILHOL, 1877), occurs in several localities at the base of the Middle Oligocene (Itardies, La Plante 2, Roqueprune 2, Soulce, Herrlingen 1). The new species C. matsoui n. sp. has been defined in the older locality of Mas de Got (top of Lower Oligocene). It is possible that the species Pseudamphimeryx decedens STEHLIN, 1910 pertains to the same genus. Also to the Tragulids must be referred the monospecific genus Iberomeryx (I. parvus GABOUNIA, 1964) from Upper Oligocene of Benara (Georgie, URSS), with which Cryptomeryx is related. These genera are not direct ancestors of Miocene tragulids; their occurrence in the Western European Oligocene results from a first immigration wawe of the family. These Tragulids are one of the most archaic groups of Ruminants. They are probably derived from a primitive stock which had acquired in Asia the selenodont condition of the dentition. 

The revision of the old collections of fossil birds from the “Phosphorites du Quercy” and the study of new material give the following results (Gruiformes, Cariamae) :  The humeri and most of the carpometacarpi described under the name Filholornis belong in Elaphrocnemus. The ulnae ascribed to Fïlholornis belong in Idiornis. Most of the post-cranial elements of the genera Elaphrocnemus and Idiornis are described and show great similarities with recent Cariamidae and Opisthocomidae, and fossil Bathornithinae.  A new genus and a new species, Oblitavis insolitus, are created in the sub-family Idiornithinae; two new species are described in the genera Elaphrocnemus (E. brodkorbz) and Idiornis (I. itardiensis), and the species Elaphrocnemus gracilis is transferred to the genus Idiornis.  The genus Propelargus Lydekker is transferred from the family Ciconiidae to Cariamidae.  A new generic name, Occitaniavis, is created for the species Geranopsis elatus, which belong in Cariamidae, while the type-species of the genus, Geranopsis hastingsiae, is a member of the Gruidae.  The affinities between the Quercy avifauna and the Neotropical one is emphasized by the occurrence of Phorusrhacidae, previously known only from the Cenozoic of South America and the Late Pliocene or Early Pleistocene of North America. Thanks to the material collected during the new excavations, the stratigraphical position of most of the species is stated precisely, and evolutionary lineages are outlined. This study shows that the suborder Cariamae, presently restricted to two South American genera, was already extremely diversified during the Eocene, and widespread in Europe and North America. 

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

A skull of Parabos cordieri, undamaged by fossilisation, allows us to make an endocranial cast, which is compared to those of Bos taurus, Boselaphus tragocamelus and Hippotragus equinus. These are studied by general form (shape, volume, « rolling-up›› and inclination), and by detailed morphology (gyri and sulci patterns, rete mirabile, importance of rhinencephalon and cerebellum).
A preliminary study is made in order to recognize convolution topography, gyri, sulci, and the various encephalic regions as clearly on an endocranial cast as on a bare encephalon; an intermediate stage is the study of an encephalon covered with «soft» meninges.
The comparative study reveals great similarity between the endocranial cast of Parabos cordieri and that of Boselaphus tragocamelus, both belonging to the tribe Boselaphini. On the other hand, a marked differences separates the cast of Parabos cordieri from that of Bos taurus, representing the tribe Bovini to which Parabos cordieri had been referred. The endocranial cast of Hippotragus equinus, which had long been considered the descendant of Parabos cordieri, is also quite different from that of Parabos. 

A recently discovered, almost complete specimen of a hybodont tooth, allows us to describe as a new species, fairly common, but usually fragmentary teeth in the Navesink Formation of New Jersey: Hybodus novojerseyensis nov. sp.
Three additional taxa are also described : Heterodontus creamridgensis, Squalicorax sp. and Pseudocorax affinis which is noted for the first time in North America.
The whole elasmobranch fauna of the Early Maastrichtian localities of New Jersey is reevaluated and several generic assignments are changed in comparison to the list previously published in 1975 (Cappetta
& Case). 

The Squamates from Sainte-Néboule are charateristic of the Upper Eocene from the Phosphorites du Quercy. Most genera known from this locality do not survive the « Grande Coupure ». Cadurcoboa insolita gen. nov., sp. nov. is a small Boidae very characteristic of the Upper Eocene. 

Based on material from 30 localities, morphologic dental, cranial and biometric analyses have permitted the characterization of two parallel Issiodoromyine lineages, and also the definition of diverse species representing several evolutive stages. Thus it is that new lineages complete the contribution made by the Theridomyinae and Cricetidae and permit, for the Quercy in particular, additional precision in the biochronologic succession of the localities. One of the lineages is limited to the genus Pseudoltinomys LAVOCAT; the other evolves from the genus Elfomys HARTENBERGER to the genus Issiodoromys BRAVARD in GERVAIS. The latter is affected by profound anatomical changes due to a functional modification of the mastication apparatus. These changes seem to be able to be put in relation with the aridification and cooling of the climate at the end of the Eocene. At the end of the middle Oligocene, a new chewing structure is achieved. It is found in diverse living rodents that inhabit a rather arid steppe environment (Cavia, Pedetes, Ctenodactylus). To these supposed nearby ecologic conditions, these rodents have responded in a convergent fashion. It is possible to attribute to the extreme specialization of Issiodoromys its incapacity to adapt to the new climatic crisis of the end of the Oligocene. The arrival of immigrants may be considered as another cause of its disappearance at this time, complementary or not with the first. 

The wear facets of primitive rodents can be homologized with those of primitive primates and ungulates. As in primates, the jaw movement was ectental, with an increased anterior component in the lingual phase (phase ll). The buccal phase (phase I) in rodents approaches the horizontal and it tends to be reduced in importance in comparison with the lingual phase. ln more advanced rodents the efficiency of grinding is increased by the development of additional cutting edges of enamel (e.g. enlargement of hypocone, development of mesoloph and lingual sinus). The buccal phase movement becomes lined up with the lingual phase movement to form a single oblique chewing stroke,resulting in planation of the crown. As the stroke becomes more longitudinal (propalinal) the enamel edges become more transverse. In Muridae propalinal chewing evolved before the loss of cusps, facets were reorientated and additional cusps developed. 

Is hereby described an impressive lchnoiauna belonging to the Lower to Middle Ludian of the Gard (S. France). The slab, already cleaned over a length of 18 m, is located near the top of the Potamides aporoschema lacustrine limestone (Lower Ludian, Euzet zone). It is therefore older than the Célas sandstone deposit, and still more than the Melanoides albigensis and M. acutus marly limestone corresponding to the Upper Levels of the Ludian stage. Although biostratigraphically older than the La Débruge and Montmartre zone, the biotope shows already a sampling of very tall Artiodactyles, Perissodactyles and Carnivorous. One of the most « majestic ›› Artiodactyles, Anopolotheriipus lavocati, nov., points out a huge size type. To mention also among the Ichnotypes described, 10, the big Perissodactyle Palaeotheriipus similimedius, nov., and the big Carnivorous Hyaenodontipus praedator, nov. 

The late Pliocene site (MN 16) of Balaruc II (Hérault, France) has provided remains of the following amphibians and reptiles: Chelotriton pliocenicus nov. sp. and Triturus marmoratus (Salamandridae), cf. Rana (Ranidae), cf. Blanus (Amphisbaenidae), cf. Agama (Agamidae),  Gekkonidae indet., Lacerta s.l. (Lacertidae), "Ophisaurus" sp. (Anguidae), Michauxophis occitanus (Aniliidae), Erycinae indet. (Boidae), Elaphe cf. E. longissima and Malpolon sp. (Colubridae), cf. Naja (Elapidae) and Vipera sp. (Viperidae). The salamandrid Chelotriton pliocenicus and the aniliid Michauxophis occitanus constitute, up to now, the only records of these groups in the European Pliocene. The fauna is indicative of a warm, dry
subtropical climate. 

The Montredon site has yielded about hundred rhinoceros remains:
- twenty two of them, including 14 carpal and tarsal bones and 6 complete metapodials, belong to
Dicerorhinus schleiermacheri at its second evolutionary stage;
- fifty one remains including a nearby complete but crushed skull, a mandible, 26 isolated cheek-teeth, 10 carpals and tarsals, one metacarpal, are of Aceratherium incisivum, second evolutionary
stage;
- fifteen remains belong to Aceratherium (Alicornops) simorrense (among other an upper molar, 8 carpals and tarsals, one metatarsal);
- six remains are attributed to an undetermined species of what is probably the most recent Prosantorhinus ever found.
The evolution stages of the two first species allow us to date the deposit back to the Upper Vallesian, MN 10 zone; Montredon is one of the youngest sites in which Aceratherium simorrense was found. The four rhino species indicate a swampy forest biotope 

Le Mémoire Extraordinaire 1988 de PALAEOVERTEBRATA regroupe dix articles consacrés au gisement à mammifères du Miocène supérieur de Montredon (Hérault), connu et classique depuis la fin du siècle dernier, et auquel est lié le nom du savant paléontologue lyonnais Charles Depéret.
Cette monographie vient normalement à la suite de celle parue en 1982 dans PALAEOVERTEBRATA, dont les différents articles traitaient de la stratigraphie du gisement, et faisaient l'étude des différents groupes de micromammifères représentés dans la faune (insectivores, chiroptères, rongeurs). 

Several endemic deer remains from the Middle Pleistocene deposits of the Castiglione cave (Oletta, Haute-Corse) are examined here. A morphometric analysis allows to relate them to a new insular subspecies Cervus elaphus rossii. The bones were compared with those of the mainland early Middle Pleistocene subspecies Cervus elaphus acoronatus Beninde and the European species Cervus elaphus Linné (Late Middle Pleistocene and Upper Pleistocene forms (continental and insular)). The Castiglione fossil shows peculiar morphofunctional features in its appendicular skeleton suggesting a morphological convergence with certain Bovidae. 

The middle Palaeocene of São José de ltaboraí (State of Rio de Janeiro, Brazil) has produced a rich and diverse fauna of boid snakes. It comprises six or seven species: Hechtophis austrinus gen. et sp. nov., Corallus priscus sp. nov., Waincophís pressulus sp. nov., Waincophis cameratus sp. nov.,"Boinae A", and "Boinae B". Moreover, two dentaries might pertain to either H. austrinus or "Boinae B", or even represent a distinct taxon. Hechtophis austrinus is assigned, with reservation, to the Erycinae. All other taxa are referred to the Boinae. The vertebrae of all taxa have paracotylar foramina, which raises the problem of the apomorphic or plesiomorphic nature of this feature. This fauna also raises the question of the presence of extinct erycine boids in South America, but it does not allow this question to be settled. 

The East African Miocene Macroscelidea, lnsectivora and Chiroptera are revised on the basis of new material. New taxa proposed are: Miorhynchocyon, .n. gen. (Macroscelididae): Míorhynchocyon meswae, n. sp.: Pronasílío ternanensis. n. gen.. n. sp. (Macroscelididae); Hiwegicyon juvenalis, n. gen. n. sp. (Macroscelididae); Parageogale, n. gen. (Tenrecidae): Prochrysochlorinae, n. subfam. (Chrysochloridae): Propottininae, n. subfam, (Pteropodidae); Chamtwaria pickfordi, n. gen., n. sp. (Vespertilionidae). Gymnurechnínus songhorensis is synonymised with G. camptolophus. The new material provides additional information on the dentition, especially of Myohyrax oswaldi. Galerix africanus. Amphechínus rusingensis, Protenrec tricuspis and Parageogale aletris. Partial skulls are described of Amphechinus rusingensis, Protenrec tricuspis, Prochrysochloris míocaenicus and Taphozous incognita. The oldest member of the Macroscelidinae (Pronasilio) is described from Fort Ternan. Galerix africanus is closely related to G. exilis from Europe. Amphechinus rusingenesis is compared with Asiatic Oligocene Erinaceinae. The Miocene age of Crocidura is rejected. On the evidence of humeri, the following families of Chiroptera are newly reported: Pteropodidae, Nycterididae, Vespertilionidae, Molossidae. Propotto is regarded as an offshoot from the Pteropodidae, not ancestral to modern forms. Chamtwaria is a primitive vespertilionoid, provisionally placed in the Kerivoulinae. Erinaceidae probably entered Africa at the beginning of the Miocene, before 20 Ma. Faunistic differences between deposits are largely to be ascribed to differences in local environment. 

The study of the bovids from Al Jadidah (Hofuf Formation, Saudi Arabia) confirms that the fauna comes from a pre-Hipparion level. The Al Jadidah age is close to that of Fort Ternan (14 m.y.) and Beni Mellal, but cannot be older than that of Fort Ternan. The age of the Hofuf Formation is close to but slightly older than the oldest deposits of the Ngorora Formation (Kenya). 7 to 9 species have been recorded, of which 2 to 4 remain indeterminate. If the great specific diversity of te bovids from this locality gives evidence of immigrations from anterior Asia (Turkey) (e.g. Pachytragus Iigabuei sp. nov.), the bovid assemblage of Al Jadidah results in fact from a double influence: from the anterior Asia and mainly from Africa (e.g. the Caprotragoides lineage and the Neotragini? Homoiodorcas). The Al Jadidah bovids reflect, on the whole, the predominant character of open to very open environment, which supports the conclusions drawn from our two preliminary studies. 

Abundant crocodilian fossil remains have been collected, mainly by R. Lavocat. from several levels of the
Palaeogene of the Tilemsi valley (eastern Mali). Several dyrosaurids occur in the Palaeocene (Montian ?) : Phosphatosaurus sp., Hyposaurus bequaerti (Dollo). H. wilsoni (Swinton). H. nopesai (Swinton), Rhabdognathus rarus Swinton, R. compressus n. sp. ln the lower Eocene has been found a dyrosaurid which cannot be accurutely determined.In the middle Eocene were found a dyrosaurid, Tilemsisuchus lavocati n. g., n. sp., and an indeterminate eusuchian. 

The present research deals with the remains of a lagomorph found at Valdemino cave and comes to the conclusion that it is a rabbit with peculiar characteristics in comparison with the other known species Oryctolagus laynensis, O. lacosti and 0. cuniculus. We studied other fossil remains of rabbit populations from Villafranchían and middle Pleistocene deposits and compared them with data from the literature and with recent material. The analysis leads us to maintain two phylogenetic hypotheses about the history of Oryctolagini. The ñrst one, already formulated by Lopez Martinez, suggests that 0. cuniculus derives from O. laynensis,while the origin of O. lacostí is unknown; according to the second hypothesis 0. laynensis would be the common ancestor of two phyletic lineages, 0. lacosti and 0. cuniculus. In both cases the lagomorph from Valdemino would be the form derived from 0. lacosti, from which however it differs in peculiar characteristics. Since the rabbit from Valdemino survives until the beginning of Postgalerian, its disappearance may coincide with the retreat of 0. cuniculus from western Europe in Spain and, perhaps, in south-western France, before the last glaciation. O. cuniculus survived in Spain, from where it spread once again over western Europe as a result of man.

  

Revision of ancient collections and study of new material from Provenchères-sur-Meuse (Rhaetian) lead to signíficant changes in the faunal list of this site. This bring to us important information about the effect of the rhaetian transgression on the evolution of the faunas at this period of time. However, study of
vertebrate microremains is always difficult and some points remain obscure, like the origin of the prosauropods which yield some remains at Provenchères-sur-Meuse and the survival in Europe during the Rhaetian of temnospondyl arnphibians different from the Plagiosauridae.