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Octodontid-like Echimyidae (Rodentia) : an upper Miocene episode in the radiation of the family
Diego H. Verzi, Maria G. Vucetich and Claudia I. Montalvo
Keywords: Argentina; Echimyidae; Miocene; New taxa; Rodentia; South America
 
  Abstract

    Reigechimys octodontiformis gen. et sp. novo and R. plesiodon sp. novo are described. They represent the frrst record of the family Echimyidae for the Cerro Azul Formation (Huayquerian Age, Late Upper Miocene) at La Pampa Province, central Argentina. Both species have hypsodont cheek teeth with an eight-shaped occlusal design. This dental morphology represents a noticeable case of convergence to octodontids and indicates that these echimyids inhabited open environments. 


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Published in Vol. 23, Fasc. 1-4 (1994)

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Hexanchiforme nouveau (Neoselachii) du Crétacé inférieur du Sud de la France
Henri Cappetta
Keywords: Hexanchiformes; New genera; Southern France; Systematics; Valanginian
 
  Abstract

    The dentition of Welcommia bodeuri nov. gen. nov. sp. from the Valanginian of Southem France is described and reconstructed. Species and genera of Upper Jurassic and Lower Cretaceous Hexanchiformes are reviewed and discussed.
    The genus Notidanoides MAISEY, 1986 must be restricted to the single Nusplingen Upper Jurassic specimen, whose attribution to the species muensteri AGASSIZ, 1843 remains doubtful.
    The genus Paranotidanus WARD &THIES, 1987 that does not rest on any type-species nor on any precise dental characterization must be rejected.
    The genus Eonotidanus PFEIL, 1983, based on a very poorly preserved and heterogeneous type-material must also be rejected.
    Teeth from the Lower Cretaceous, with a peculiar morphology, previously assigned to Eonotidanus or to Notidanoides, are to be ranked in the new genus Pachyhexanchus


  Article infos

Published in Vol. 20, Fasc. 1 (1990)

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Revision des faunes de vertébrés du site de Provenchères-sur-Meuse (Trias terminal, Nord-Est de la France)
Gilles Cuny
Keywords: amphibians; Fishes; Reptiles; Rhetian; Triassic
 
  Abstract

    Revision of ancient collections and study of new material from Provenchères-sur-Meuse (Rhaetian) lead to signíficant changes in the faunal list of this site. This bring to us important information about the effect of the rhaetian transgression on the evolution of the faunas at this period of time. However, study of
    vertebrate microremains is always difficult and some points remain obscure, like the origin of the prosauropods which yield some remains at Provenchères-sur-Meuse and the survival in Europe during the Rhaetian of temnospondyl arnphibians different from the Plagiosauridae.

      


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Published in Vol. 24, Fasc. 1-2 (1995)

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Fossil mammals and the age of the changxindian formation, Northeastern China
Spencer G. Lucas
Keywords: Changxindian Formation; China; Eocene; Fossil mammals
 
  Abstract

    Re-evaluation of the small collection of mammal fossils from the Changxindian Formation near Beijing, China indicates the following taxa are present: Eutheria, Hypsimilus beifingensis, cf. Miacis sp., Anthracotheriidae and Forstercooperia grandis. The presence of Forstercooperia grandis indicates an Irdinmanhan age and does not support previous assignment of a Sharamurunian age to the Changxindian Formation. 


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Published in Vol. 25, Fasc. 2-4 (1996)

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The lower Miocene artiodactyls of Tagai bay, Olhon island, lake Baikal (Russia)
Inesa Vislobokova
Keywords: Artiodactyls; Mammals; Miocene; Siberia
 
  Abstract

    The mammalian fauna of the Tagai locality of Olhon Island in Lake Baikal contains six artiodactyls:
    Amphitragulus boulangeri, Lagomeryx parvulus, Stephanocemas sp., Palaeomeryx kaupi, Orygotherium aff. escheri, and Brachyodus intermedius. They are reported from Eastern Siberia for the first time. A description of teeth and limb-bones are given. An analysis of the systematic composition of the Tagai fauna allows it to be correlated with European faunas of the second part of the Early Miocene (Orleanian, MN 3-4) and permit suggestion of the existance of a united European-Siberian zoogeographical sub-area in the middle latitudes of Eurasia at that time. 


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Published in Vol. 23, Fasc. 1-4 (1994)

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Batoids (Rajiformes, Torpediniformes, Myliobatiformes) from the Sülstorf Beds (Chattian, Late Oligocene) of Mecklenburg, northeastern Germany: a revision and description of three new species
Thomas Reinecke
Keywords: Batoids; Chattian; Elasmobranchii; North Sea Basin; Oligocene

doi: 10.18563/pv.39.2.e2
 
  Abstract

    Bulk-sampling of fossil-rich tempestites from the Chattian Sülstorf Beds of
    Mecklenburg, north-eastern Germany, yielded a rich selachian fauna in which batoids
    predominate by the abundance of teeth but are subordinate by the number of taxa. Thirteen
    taxa are identified, among which rajiform batoids are the most diverse (six species). One
    genus and three species are newly described: Raja thiedei sp. nov., Oligoraja pristina gen. et
    sp. nov., and Torpedo chattica sp. nov. Two species are reallocated: Atlantoraja cecilae
    (Steurbaut & Herman, 1978) new comb., and Dipturus casieri (Steurbaut & Herman, 1978)
    new comb. Ontogenetic heterodonty is documented for the first time in the dental pattern of
    Myliobatis sp. Stratigraphical ranges of batoid taxa in the period from Rupelian to Langhian
    are presented and partly discussed in context with the palaeoclimatic evolution and
    palaeogeographic situation of the North Sea Basin. 


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Published in Vol.39-2 (2015)

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A late Eocene palaeoamasiine embrithopod (Mammalia, Afrotheria) from the Adriatic realm (Island of Rab, Croatia)
Fabrice Lihoreau, Ljerka Marjanac, Tihomir Marjanac, Ozan Erdal and Pierre-Olivier Antoine
Keywords: Balkanatolia; Grande Coupure; Great Adria; Paleobiogeography; Systematics

doi: 10.18563/pv.47.1.e1
 
  Abstract

    A cheek tooth recently unearthed in the Lopar Sandstone unit, of late Eocene age, in the northern part of Rab Island, Croatia, is one of the very few Eocene mammalian remains found in the Adriatic area. Thorough comparison of this tooth with those of Old-World Palaeogene mammalian orders suggests that it is a M3 belonging to an embrithopod afrothere. The specimen is referred to as Palaeoamasia sp. This genus was formerly known only in Eocene deposits of Anatolia but with close relatives in Romania among Palaeoamasiinae. The geographical distribution of this subfamily perfectly matches the recently-named Balkanatolian landmass, which experienced in-situ evolution of endemic mammals prior to the Grande Coupure event that occurred around the Eocene–Oligocene transition. This last event is characterised by massive Asian immigration in Western Europe and the supposed extinction of many endemic Central and Western European mammals, including Palaeoamasiinae.
      


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Published in 47-1 (2024)

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New remains of the very small cuckoo, Chambicuculus pusillus (Aves, Cuculiformes, Cuculidae) from the late Early/early Middle Eocene of Djebel Chambi, Tunisia
Cécile Mourer-Chauviré, El M. Essid, Hayet Khayati Ammar, Laurent Marivaux, Wissem Marzougui, Rim Temani, Monique Vianey-Liaud and Rodolphe Tabuce
Keywords: Cuckoos; Eocene; Fossil bird

doi: 10.18563/pv.40.1.e2
 
  Abstract

    Abstract: A very tiny cuckoo, Chambicuculus pusillus, was recently described after a few fragments of tarsometatarsi from the late Early/early Middle Eocene of Djebel Chambi, Tunisia. New remains, notably a coracoid, confirm the attribution of this genus to the recent family Cuculidae. This coracoid shows a very elongate and strap-like processus procoracoideus. This morphological feature is otherwise only known in the family Cuculidae. The characteristics of the coracoid and tarsometatarsi show that Chambicuculus is morphologically more advanced over the other stem cuculids described in Europe and North America. Chambicuculus is the oldest Cuculidae known so far. 


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Published in Vol.40-1 (2016)

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Modification du statut générique de quelques espèces de sélaciens crétacés et tertiaires
Henri Cappetta
Keywords: Cretaceous; Selachians; Tertiary
 
  Abstract

    The re-examination of six fossil selachian species has involved the creation of six new genera: Squatiscyllium, Protolamna, Parotodus, Abdounia, Physogaleus, Hypolophodon and of one new species : Prozolamna sokolovi. The modification of the generic statute of these species allows to clarify their systematic position and to define their relationships at a familial level. 


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Published in Vol. 10, Fasc. 1 (1980)

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Macroscelidea, Insectivora and Chiroptera from the Miocene of east Africa.
Percy M. Butler
Keywords: Chiroptera; East Africa; Insectivora; Macroscelidea; Miocene; Systematics
 
  Abstract

    The East African Miocene Macroscelidea, lnsectivora and Chiroptera are revised on the basis of new material. New taxa proposed are: Miorhynchocyon, .n. gen. (Macroscelididae): Míorhynchocyon meswae, n. sp.: Pronasílío ternanensis. n. gen.. n. sp. (Macroscelididae); Hiwegicyon juvenalis, n. gen. n. sp. (Macroscelididae); Parageogale, n. gen. (Tenrecidae): Prochrysochlorinae, n. subfam. (Chrysochloridae): Propottininae, n. subfam, (Pteropodidae); Chamtwaria pickfordi, n. gen., n. sp. (Vespertilionidae). Gymnurechnínus songhorensis is synonymised with G. camptolophus. The new material provides additional information on the dentition, especially of Myohyrax oswaldi. Galerix africanus. Amphechínus rusingensis, Protenrec tricuspis and Parageogale aletris. Partial skulls are described of Amphechinus rusingensis, Protenrec tricuspis, Prochrysochloris míocaenicus and Taphozous incognita. The oldest member of the Macroscelidinae (Pronasilio) is described from Fort Ternan. Galerix africanus is closely related to G. exilis from Europe. Amphechinus rusingenesis is compared with Asiatic Oligocene Erinaceinae. The Miocene age of Crocidura is rejected. On the evidence of humeri, the following families of Chiroptera are newly reported: Pteropodidae, Nycterididae, Vespertilionidae, Molossidae. Propotto is regarded as an offshoot from the Pteropodidae, not ancestral to modern forms. Chamtwaria is a primitive vespertilionoid, provisionally placed in the Kerivoulinae. Erinaceidae probably entered Africa at the beginning of the Miocene, before 20 Ma. Faunistic differences between deposits are largely to be ascribed to differences in local environment. 


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Published in Vol. 14, Fasc. 3 (1984)

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Cricetid and arvicolid rodents of the California wash local fauna, late Blancan of the san Pedro Valley, Arizona.
Cristiana Mezzabotta
Keywords: Arvicolidae; Blancan; Cenozoic; Cricetidae; Mammals
 
  Abstract

    An assemblage of micromammals is reported from California Wash, a fossil bearing continental deposit in the San Pedro Valley, Arizona, late Blancan in age. Cricetid and Arvicolid rodents are richly represented, including four and two species, respectively. This study mainly focuses on Sígmodon, the most abundant form. The sample of Sigmodon is compared to samples of the same genus from other localities of the San Pedro Valley of comparable age, and some inferences on the taxonomy of the genus are attempted. The specimens are referred to Sigmodon minor and Sigmodon cf. S. curtisi. Other cricetids (Onychomys pedroensis and Baiomys brachygnathus) and arvicolids (Mictomys vetus and Ondatra ídahoensis) are also recognized and described. 


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Published in Vol. 26, Fasc. 1-4 (1997)

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A new and primitive species of Protophiomys (Rodentia, Hystricognathi) from the late middle Eocene of Djebel el Kébar, Central Tunisia
Laurent Marivaux, El M. Essid, Wissem Marzougui, Hayet Khayati Ammar, Sylvain Adnet, Bernard Marandat, Gilles Merzeraud, Rodolphe Tabuce and Monique Vianey-Liaud
Keywords: Adaptive radiation; Bartonian; Dental morphology; North Africa; Paleobiogeography

doi: 10.18563/pv.38.1.e2
 
  Abstract

    Based on fossil discoveries and phylogenetic studies, an Eocene Asian origin for hystricognathous rodents and anthropoid primates has gained strong support in recent years. The two groups then invaded both Africa and South America, which promoted their evolutionary success. However, the fossil record has so far failed to constrain the nature and precise timing of these pivotal dispersal events. In Africa, given the apparent absence of hystricognaths and anthropoids in early to early middle Eocene localities, it is suggested that these mammal groups dispersed from Asia to Africa sometime during the middle Eocene. In this paper, we report the discovery of several isolated teeth of a rodent from a new vertebrate locality situated in central Tunisia (Djebel el Kébar, KEB-1), dating from the late middle Eocene (Bartonian, ~39.5 Myr). These fossils document a diminutive new species of Protophiomys (P. tunisiensis nov. sp.), a basal genus of hystricognathous rodents which is well known from several North African mammalian-bearing localities of the end of the Eocene. The teeth of P. tunisiensis display a suite of anatomical details comparable with those observed in the other species of the genus, but with a lesser degree of development. Such an apparent primitive evolutionary stage is corroborated by the greater antiquity of this Tunisian species. P. tunisiensis nov. sp. is so far the most ancient representative of hystricognaths in Africa. However, it can be expected that hystricognaths were already present on that landmass given the new data on early caviomorphs recently reported from South America (at ~41 Myr). The arrival of hystricognaths in Africa from South Asia certainly predates the depositional period of the Kébar sediments, but perhaps not by much time. 


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Published in Vol.38-1 (2014)

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Paleobiology of Messel Erinaceomorphs
Gerhard Storch
Keywords: Erinaceomorpha; Germany; Grube Messel; Lipotyphla; Middle Eocene; Paleobiology
 
  Abstract

    Three erinaceomorph species are known from the early Middle Eocene of Grube Messel near Darmstadt, Germany, which are referred to the family Amphilemuridae. Pholidocercus hassiacus, Macrocranion tupaiodon, and Macrocranion tenerum showed extraordinary adaptations to their different life strategies, and several of their specializations are unknown among living insectivores. Pholídocercus was a well-defended robust animal with an opportunistic feeding strategy. Macrocraníon zupaiodon was a slender forest floor-dweller with saltatorial specializations to escape from predators; fishes were the preferred component of its omnivorous diet. Macrocranion tenerum exhibited a combination of both survival strategies, extremely elongated hind limbs for rapid and even ricochetal flight and a spiny exterior as an effective protective device; it was probably specialized for feeding on ants. Thus, closely related, omnivorous-insectivorous forest floor-dwellers could exploit the Messel ecosystem. 


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Published in Vol. 25, Fasc. 2-4 (1996)

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Introduction à l'oeuvre scientifique de Donald E. Russell, "gentleman paleontologist"
Marc Godinot and Phillip D. Gingerich
Keywords: D.E.Russell; Eocene; Mammals; Paleocene; Paleontology; synthesis
 
  Abstract

    The scientific career of D.E. Russell began with a Pliocene fauna from Oregon, and then turned in the direction of European Paleogene mammals. Field work followed by study of the mammals that were collected, firstly in the Paleocene and later in the early Eocene, greatly rejuvenated learning in this field. Syntheses on the Northwest European Tertiary basin and on European marnmals and stratigraphy came next. Research on the Eocene of Asia was carried out jointly with Gingerich on Pakistan and with Dashzeveg on the faunas of Mongolia. An important synthesis on the entire Paleogene of Asia, joint with Zhai, followed. Field work in Africa with Sigogneau-Russell led to the discovery of Mesozoic mammals there. A synthesis of mammalian paleofaunas of the world was written with Savage, and a similar synthesis of Cenozoic vertebrate faunas is currently being prepared. These achievements reflect the perennial importance of field work, numerous collaborations with both amateurs and professionals, and the human qualities of this author.
      


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Published in Vol. 25, Fasc. 2-4 (1996)

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Les Chiroptères du Miocène inférieur de Bouzigues. 1- Etude systématique.
Bernard Sigé
Keywords: bats

doi: 10.18563/pv.1.3.65-133
 
  Abstract

    In recent years, the techniques of chemical preparing have permitted a rich paleontologic material to be obtained from the phosporitic sediment of Bouzigues (Hérault, France). The fauna of this locality is comprised of quite varied microvertebrates, amphibians, reptiles, birds, mammals. Twenty five species of the latter, belonging to seven orders, are today known from the site. Among them, the numerous rodents have allowed L. Thaler to chronologically situate this fauna in the Zone of Laugnac (<< late Aquitanian ›> of some authors).
    The chiropterans are, with the rodents, the best represented of the locality's mammals. Three families comprise the bat fauna, with nearly complete dominance by one of them (Hippoxideridae) over the two others (Megadermatidae and Vespertílionidae)
    Six forms are described, of which three are new species and one a new sub-genus.
    Megaderma braillomi n. sp., an animal of rather large size, shows like the Miocene megaderms several evolved dental characters, translating the adaptation of these animals to a partially carnivorous regime. The Bouzigues species seems, however, to represent a particular lineage.
    Hipposideros (Brachipposideros n. subgen.) dechaseauxi n. sp. and Hípposideros (Brachipposideros) cf. collongenris Depéret, small sized forms, belong to a group rather well represented in the late Oligocene and early Miocene of Europe, and not distinguished until now within the genus Hipposideros.
    Hipposideros (Pseudorhinolophus) bouziguensis n. sp., is the most abundant mammal in the locality and, occuring at the Oligocene-Miocene limit, the last representative known of the subgenus Pseudorhínolophus, common in Europe from the middle Eocene.
    However, beyond Neogene and Quaternary times, certain among the numerous living species of Hipposideros are close to Pseudorhinolophus and others to Brachipposíderos. 'This fact would in the future justify a global revision of the genus, on the basis of comparative anatomy of the squeleton and of the teeth.
    The bat fauna of Bouzigues is completed by two small Vespertilionidae, rare forms, Myoris sp. I and sp. II.
      


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Published in Vol. 01, Fasc. 3 (1968)

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Révision systématique des Anchilophini (Palaeotheriidae, Perissodactyla, Mammalia).
Jean-Albert Remy
Keywords: Anchilophus; Eocene; new genus; new species; Palaeotheriidae; Paranchilophus; Perissodactyla; Systematics

doi: 10.18563/pv.37.1-3.1-165
 
  Abstract

    The knowledge of the Anchilophini has been lately renewed by the discovery of a rather large amount of new material still largely unpublished. This new material offers the opportunity of a systematic revision of this tribe gathering those of European Eocene Equoidea which bear no mesostyle on upper check teeth and display a heavy trend to the molarization of premolars.

    A cladistic analysis has made out two genera, Anchilophus (Paranchilophus included as a subgenus), characterized by a marked lophodonty and the transverse narrowness of the cheek teeth, a rather high hypsodonty, the frequent occurrence of "crochets" and "anticrochets" on the superior ones, and a rather weak molarization of the premolars, opposite to Metanchilophus n. gen. whose cheek teeth are more transversally elongated, less high, less lophodont, with cusps better distinct, enamel thicker and premolars more molarized on the whole.

    Three species of Anchilophus are recognized, A. desmaresti, type species of the genus, A. (Paranchilophus) remyi and A. (Paranchilophus) jeanteli n. sp.

    The genus Metanchilophus is more diversified with the species dumasi, radegondensis, gaudini (whose a new sub-species M. g. fontensis is defined), depereli, castrensis n. sp. and chaubeti n. sp.

    The skull anatomy has been moreover described with several taxa; it brings to light (for all that one can generalize) that Anchilophini were light animals with a slender and elongated snout, a thin zygomatic arch, a rather developed encephalon with an advanced gyrencephaly.

    The structure of the nasal opening together with the occurrence of epitympanic sinuses and the molarizing process of the premolars corroborate the attribution of this tribe to the family PalaeotheIiidae. 


  Article infos

Published in Vol. 37, Fasc. 1-3 (2012)

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The Pleistocene vertebrate fauna of Robinson Cave, Overton County, Tennessee
J. E. Guilday, H. W. Hamilton and A. D. Mc Crady
Keywords: Fauna; Mammalia; Pleistocene; Tennessee

doi: 10.18563/pv.2.2.25-75
 
  Abstract

    A late Pleistocene deposit of 60 species of vertebrates and 12 of invertebrates is described from Robinson Cave, Overton County, Tennessee, U.S.A. Forty-eight species of mammals are represented by at least 2,483 individuals; 10 % are extinct, 10 % occur in the state only as boreal relicts in the Great Smoky Mountains; 23 % no longer occur as far south as Tennessee; 57 % occur at or near the site today. Nínety-one percent of the Recent mammal species can be found living today in the Minnesota-Wisconsin area, approximately 10 degrees farther north. Fluorine analysis suggests a long period of accumulation. The following 10 mammalian species are recorded from Tennessee for the first time. Sorex arcticus, Microsorex hoyi, Citellus tridecemlineatus, Clethrionomys gapperi, Microtus pennsylvanicus, Synaptomys cooperi, Synaptomys borealis, Zapus nudsonius, Napaeozapus insignis, Martes americana. Six additional species are present as boreal relicts in the Great Smoky Mountains of eastern Tennessee but not at the site today : Sorex cinereus, Sorex dispar, Sorex palustris, Parascalops breweri, Glaucomys sabrinus, Mustela nivalis. Six forms are extinct: Canis dirus, Ursus americanus amplidens, Sangamona furtiva, Dasypus bellus, Mammut americanus,Megalonyx jeffersoni. Twenty-six additional species of mammals, all of the snails, birds, reptiles, and amphibians recovered from the fauna still inhabit the area today: The fauna is indicative of a cold-temperate climatic episode associated with the Wisconsin glaciation, but may be chronologically mixed. 


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Published in Vol. 02, Fasc. 2 (1969)

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Evolution of the Rhizomyine zygoma
Lawrence J. Flynn, Mohammed Sarwar and Jean-Jacques Jaeger
Keywords: parallel evolution; Rhizomyidae; Rodentia; Siwalik; zygoma
 
  Abstract

    Cranial anatomy of a late Miocene rhizomyid, Brafhyrhizomys cf. B. pilgrimi, provides new evidence on the origin of the dorsal, round infraorbital foramen of living rhizomyines. Primitive rhizomyids retain a myomorphous keyhole foramen with a long ventral slit that retracts upward during the evolutionary history of the Rhizomyidae. The primitive condition of the elongated ventral slit is represented by Kanisamys sivalensis. Among later burrowers the foramen shows progressive dorsal migration, the ventral slit terminating midway up the snout in B.tertracharax and B. choristos ; well above the midline of the snout in Brachyrhizamys cf. B. pilgrimi. Apparently this transformation began earlier among Rhizomyinae than among Tachyoryctinae and continued to a more derived stage in rhizomyines. ln living Rhizomyx the ventral slit is absent and only a high round hole remains at the anterior end of the zygomatic arch. 


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Published in Vol. 15, Fasc. 3 (1985)

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Study of the Turolian hipparions of the lower Axios valley (Macedonia, Greece). 4. Localities of Dytiko.
George D. Koufos
Keywords: Equidae; Greece; Hipparion; Lower Axios Valley; Macedonia; Mammalia; Turolian
 
  Abstract

    The hipparions from the Dytiko localities of the lower Axios valley (Macedonia, Greece) are studied. The material comes from three localities Dytiko-l, 2, 3 (DTK, DIT, DKO), which are situated near the village of Dytiko, about 60 km northwest to Thessaloniki. Three species have been determined, the medium-sized H. mediterraneum, the small-sized H. matthewi and the very small-sized H. periafricanum. The determined Hipparion species, their morphological characters and their comparison with the other Axios valley material indicate a Late Turolian age for the Dytiko localities. 


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Published in Vol. 18, Fasc. 4 (1988)

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Lissamphibians from Dams (Quercy, SW France): Taxonomic identification and evolution across the Eocene-Oligocene transition
Alfred Lemierre and Maeva J. Orliac
Keywords: Eocene-Oligocene; Grande Coupure; Lissamphibia; Quercy Phosphorites

doi: 10.18563/pv.48.1.e3
 
  Abstract

    The locality of Dams (Quercy, southwestern France) has yielded two fossil assemblages, one from the late Eocene and another from the early Oligocene, making it one of the few localities with infillings across the Eocene-Oligocene transition. At least 24 taxa (13 mammals, 11 snakes) have been identified in this locality. Study of the lissamphibian remains from Dams yields an Eocene and an Oligocene assemblage, with a total of eight taxa. The Eocene assemblage includes two unnamed salamandrine species, one unnamed pelobatid species and one pyxicephalid species (Thaumastosaurus). The Oligocene assemblage includes two unnamed pleurodeline species, one salamandrine species (Salamandra sansaniensis) and an unnamed pelobatid species. Among the eight taxa from Dams, one Eocene salamandrine and one Oligocene pleurodeline are identified for the first time in the Quercy. A review of the lissamphibians from the Quercy area identifies eleven taxa for the Late Eocene (MP19) and eight taxa for Early Oligocene (MP22), with a major turnover at the Eocene-Oligocene transition. This turnover occurs in a time of major climatic changes, with a significant decrease in temperature and precipitation and concurrent increase in seasonality in Europe, likely affecting specialized taxa.
      


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