The Palaeobatrachidae until now have not been reported from France. The present note makes known the discovery of very significant remains in three Tertiary localities of France: Cernay (late Paleocene), Laugnac (late Aquitanian) and Sansan (middle Helvetian). The osteologic characters, as well as the geographic and stratigraphic distribution of the family are re-stated with added details. The Palaeobatrachidae are a European family known from the late Jurassic of Catalonia (publication in course by J. Seiffert) to the Plio-Pleistocene (pre-Mindel) of Poland and Rumania. By their way of life and certain osteologic characters they are similar to the Pipidae, but they are distinguished by profound differences. It would be rash to include them in the Pipoidea or even in the Aglossa. They are apparently an independant group, essentially holarctic, which held in Europe a role equivalent to that played by the Pipidae south of Tethys. 

The first lists of Insectivores (Erinaceidae, Talpidae and Soricidae) from the Pliocene beds of Southern France and North-East Spain are given in this paper. The material from twelve localities is studied. These localities are geographically situated in Languedoc (Celleneuve, Vendargues, Nîmes, Sète, Balaruc 2 and Seynes), in Roussillon (Terrats, Serrats-d'en-Vacquer, Château d'eau and Mont-Hélène) and in North-East Spain (Layna, Medas Islands and Puebla de Valverde). These faunas correspond to the Early, Middle and Late Pliocene. 1 to 8 taxa are identified in these localities and 14 specific taxa are presently listed for this period in this area. Two new specific taxa are described as Galerix depereti nov. sp. from all the Early Pliocene localities in the North-Pyrenean area and as Desmanella gardiolensis nov. sp. from Balaruc 2. For this small mammals, two faunal assemblages are recognized. The first one is dated from the Early Pliocene (F 1, 2 and 3 zones in Aguilar et Michaux) and is characterized by Galerix depereti and rare and little diversified Soricids. The second one is Late Pliocene in age (zones G 2 and G 3). The fossils of the genus Talpa are relatively abundant and the Soricids are diversified and very abundant. The Middle Pliocene (zone G 1) is a transitional period. ln these faunas, most of the insectivore genera are known from the European Late Miocene beds (8 on 10). This fact demonstrates a relative continuity between the invectivore faunas from the Late Miocene to the Early Pliocene. In conclusion, somme paleoecological considerations are suggested.
  

New Dendromus species (Rodentia, Muroídea) from Langebaanweg (Pliocene, South Africa). Stratigraphical and paleoecological consequences.

Two new species of Dendromus are described from the Langebaanweg site which precises the evolutionary trend among this genus in South Africa and gives further paleoenvironmental indications. Two evolutionary stages are described: D. darti nov. sp. shows low-crowned molars with bunodont cusps and its more closest relative would be D. melanozis from the Cape region. On the contrary, D. averyi nov. sp. is more lophodont and is better related with the modem D. melanotis. Both species are at a less evolved stage than the Dendromus sp. from Laetolil Beds at Laetoli. The Langebaanweg deposits cannot still be dated by biostratigraphy but they clearly cannot be older than the basis of Pliocene times. The association of Dendromus and Mystromys in the same levels indicates a grassland environment with woodland patches as well as probable swamps. 

A well-preserved anterior cranium of Paranourosorex gigas RZEBIK-KOWALSKA, 1975 from Podlesice, Poland, provides additional knowledge of this rare shrew, including almost complete maxillary dentition. The systematic status of this species as a relative of recent Anourosorex MILNE-
EDWARDS, 1870 is confirmed 

     The Faunas of Rodents from seven north-african fossiliferous beds distributed from the Middle up to the Uppest Miocene are studied. One genus, seventeen species, one subspecies described are new.
       A detailed description is given of the evolution of the numerous specific lines, systematic groups and communities. The evidence of the strong endemism of the North African faunas of that time compared with those of Europe as well as of Tropical Africa is clearly shown. Special relationships with Asia clearly also appear, mostly in the Middle Miocene and the beginning of the Upper Miocene.
       Using the specific lines with a biostratigraphic purpose gives the opportunity to establish a biochronological scale for the continental formations of the Maghreb. This scale is calibrated by a few absolute datations. These datations, as well as the stratigraphical study of several fossiliferous localities have given the possibility to establish precise correlations with the corresponding scales of Europe and of Tropical Africa, as well as with the marine formations of the Neogene of occidental Mediterranean sea.
       The paleoclimatic data connected with this period in North Africa are explained and the contribution of the micromammals is discussed. 

La localité fossilifère du Puech de Montredon, désignée plus communément sous le nom de Montredon, est située sur la commune de Montouliers (Hérault) à quelques 300 mètres de la limite avec le département de l'Aude. Elle a été découverte en 1845 par Narbonne, Directeur des Mines de La Caunette, et de très nombreux restes de vertébrés continentaux y ont été extraits. La plus ancienne mention de ce gisement dans la littérature semble être celle de Lartet (1859) qui signale que "M. Jourdan, de Lyon, a constaté à Montredon, près de Bize (Aude), l'association des restes de Dinotherium avec l'Hipparion". 

Six orders of birds adapted to aquatic life are represented among the numerous avifauna of "Saint-Gérand-le-Puy": Gaviiformes, Procellariiformes, Pelecaniformes, Ciconiiformes, Phoenicopteriformes, and Anseriformes. The present study of this avifauna proposes several changes in systematics:- Procellariiformes: Puffinus arvernensis does not belong in Procellariidae but in Diomodeidae, and it is transferred to the fossil genus Plotornis previously described in the Middle Miocene of France. - Pelecaniformes: Phalacrocorax littoralis remains in Phalacrocoracidae; P. míocaenus is different from the modern species, and is transferred to the new genus Nectornis. Empheresula arvernensis, described in the Oligocene deposits of Gannat, seems to be present in Saint-Gérand-le-Puy too. Pelecanus gracilis shows many differences from the modern species, and belongs to the new genus Miopelecanus, - Ciconiiformes: Ardea formosa nom. oblit. is a synonym of Proardeola walkeri. - Anseriformes: a new species closely related to swans is described, and belongs to the fossil genus Cygnopterus, of the Middle Oligocene of Europe; this species is called C. alphonsi. The ecology of each species is suggested by comparison with that of its nearest living relatives, and by study of osteological adaptations. 

Near "la Lieude", in the Lodève basin, more than a thousand of footprints are distributed in a twenty of trackways which amounts to 220 m length. They have been found on calcareous siltstone level in the B site named also "Réserve Naturelle Volontaire". This last is located in the Saxonian summit dated Upper Permian. "La Lieude" tracks are described by using statistical methods then they are compared with others from the world Permian. What allows to distinguish 4 following ichnotaxa: Lunaepes ollierorum nov.ichnosp., Merifontichnus thalerius nov. ichnogen. and nov. ichnosp., Planipes brachydactylus nov.ichnosp. and Brontopus circagiganteus nov. ichnosp. All these traces are attributed with possibility or probability to Therapsida or to Therosauria, except Brontopus circagiganteus nov. ichnosp. that could be due to Caseamorpha. All these animals whose sizes have been estimated between l and 5 m lived probably in a playa environment.The biological and sedimentological data from "la Lieude" footprints levels compared with informations provided by the tracks orientations, suggest the following scenario. Animals coming from the North have crossed a sandy channel bank with plants zones by directing to the South for the majority. Maybe, they were going to the lacustrine part of the playa, close to "la Lieude" footprints that they have just trampled on. 

Dichobunoid artiodactyls are described in this paper from the middle Eocene Geiseltal lignite deposits near Halle (Sachsen-Anhalt, Germany). The genera Eurodexis and Parahexacodus are established based on odontologica1 studies. The type-species are E. ceciliensis (FRANZEN & KRUMBIEGEL, 1981) and P. germanicus n. sp. from the "obere Mittelkohle" (see text), MP 13. The genera are referred to the new subfamily Eurodexeinae (Dichobunidae, Artiodactyla, Mammalia), which also contains the monospecific genus Eygalayodon SUDRE & MARANDAT, 1993. These new genera show many features similar to North American homacodontids and antiacodontids. They exemplify a high degree of diversity in European dichobunids. Moreover, the family Dichobunidae includes the Dichobuninae and the Hyperdichobuninae. The connection of the posthypocristid with the postentocristid, the shift of a medial to a lingual position of the hypoconulid on the lower molars and the development of a large, caniniform P1 are regarded as the principal tendencies of the eurodexeines. The entoconid is larger and more mesially situated compared to that of Diacodexis. The hypoconulid has a lingual position on a broad postcingulid. The elongation of the premolars and the presence of diastemata are considered associated with extension of the muzzle. This and the acute tubercles of the molars could indicate a limited degree of insectivory. Protodichobune cf. oweni and Diacodexis cf. varleli, first reported here from the "untere Unterkohle" (MP 11) of the Geiseltal, demonstrate the existence of forms in the basal part of the middle Eocene, that are direct descendants of lower Eocene ones. Eurodexis russelli n. sp. from MP 10 of Premontre is the most primitive representative of Eurodexis. A further part of this lineage could be Messelobunodon sp. from MP 11 of Messel (Germany). Another form from Premontre, referred to Eurodexeinae indet., represents the origin of the lineage to Parahexacodus germanicus n. sp. from the Geiseltal (MP 13). The identification of several lineages in MP 10 is evidence of an adaptive radiation during this time. Unfortunately the history of the earlier species of Diacodexis is not known well enough to exclude earlier diversifications. Our study shows the independent development of artiodactyls at the end of the lower Eocene in Europe, North America (and maybe in Asia) with much convergence. The question of the geographic origin of the genus Diacodexis is still open. 

The fauna studied in the following work involves the dentitions and skulls more or less complete of 5 genera, among which only Cephalomys was previously known by its skull. One must notice that the Salla's species of this genus is a new one. Sallamys, rather small, shows a dentition rather similar to that of Platypittamys Wood from Patagonia. The upper molars, more primitive than those of this last genus, according to the smaller dimensions of the hypocone, retain a distinct metaloph. This metaloph tends to be reduced in a way which may give us a possibility to understand how it disappeared in Platypittamys. The upper P4 can be compared as well to that of Platypittamys as to that of Gaudeamus from the African Oligocene. The lower P4, more molarized than that of Platypittamys, is already moving towards the miocene type of structure. The infraorbital foramen is wide and the insertion of the masseter on the muzzle is spacious. Branisamys, genus of a great size, shows an auditory region partly preserved, peculiarly the promontorium with the fenestra rotunda, entirely of the Hystricognathi type. Upper molars are very clearly pentalophodont. A new reconstruction is proposed for the tooth called Villarroelomys by Hartenberger. This tooth is shown to be a lower D4, perhaps of Branisamys , certainly of a rather nearly allied form, and Hartenberger does agree with the essential part of this new conclusion. Of Incamys, two incomplete skulls are known, each one being admitted to be the type of a distinct species, the first one being I. bolivianus, I. pretiosus the second. The infraorbital foramen is of a great size and the impression of the masseter on the muzzle is spacious. The sphenopalatine foramen is widely developed and of a really very uncommon great size. Only Thryonomys from Africa shows a similar tendency to the enlargement of this foramen, but not so extreme. The main basicranial foramina can be observed. The upper teeth, hemi-hypsodont, show, either a vestigial metaloph, similar to that of recent Thryonomys from Africa, associated with a well developed mesoloph, either a well developed metaloph, while the mesoloph is reduced or absent. Cephalomys was previously known by anterior parts of the skull showing a wide infraorbital foramen and a spacious facial insertion of the masseter. Its lacrymal is of the phiomorph type and the spheno-palatine foramen is seemingly of great size, like in Incamys. The species is new. The varied peculiarities of the upper teeth of these genera can be easily understood if we refer to the plan of the teeth of Phiomys andrewsi from the Oligocene and Miocene of Africa. The structure of this genus, clearly more primitive, still typically brachyodont, shows and clearly explains the fundamental coherence of the varied realisations arised from such a structure. Luribayomys n.g. is known only by an anterior half of a skull without teeth. It is remarquable by the great development of the masseter's insertions on the muzzle and by the lacrymal region, well preserved, typically phiomorphid. The classification previously published by A.E. Wood and B. Patterson is granted in its essential parts, provisionally, but not as a definitive solution. Nevertheless the Dasyproctidae are integrated within the Cavioidea, following the conclusions of Bugge and of Vucetich, reached independently. The conclusion emphasizes the exceptional meaning of the fauna of Salla-Luribay. This shows that Platypittamys, while interesting, can no more be supposed certainly representative of the normal structure of the Oligocene Caviomorph, and not even of their ancestors. The anatomical peculiarities exhibited in these new samples, auditory region, lacrymal, spheno-palatine foramen, reinforce the primitive structural identity with the Phiomorpha. Similarly, the new lower D4 favour very close relationships, ever if the affinities of the D4 has been questioned or minimized by Wood and Patterson. It is certainly possible to admit that parallelism could explain limited similarities, like the presence in North America of Rodents with an hystricomorph type of infraorbital foramen and an hystricognath mandible. But if the parallelism could be a sufficient explanation of the identical association of multiple and complete structures observed in the Caviomorpha and Phiomorpha, all the Zoological systematic would have to be questioned. The last positions of A.E. Wood on the subject (1975) are revised and criticised, and the recent publications studying the problems of distance between Africa and South America in Eocene time, as a consequence of the drift, are quoted; the possibility of transportation by rafts is shown. A new hypothesis is proposed about the interrelationships of Pentalophodont Rodents, with interesting paleobiogeographic implications. 

A recent campaign of excavations (1965-68) undertaken by the Laboratoire de Paléontologie of Montpellier in pockets of the Quercy phosphorites, has permitted the dating of several localities thanks to the analysis of their micromammalian fauna.
The rodents of localities belonging to the middle Oligocene (La Sauvetat zone) are the object of this study. This rodent fauna has been compared to the forms coming from three stratified localities belonging to the biochronologic zone of La Sauvetat :  Balm, Montalban and Lovagny.
The abundance of material, notably of theridomyids and cricetids, permitted a summary statistical study to be effected and gave some precise details on the variability of the species (Theridomys varians in particular). Other details were provided on certain groups, notably on the upper teeth of Scíuromys cayluxi and on the milk teeth of theridomyids. Some specimens of «Sciurus» sp. and of Plesispermophilus angustidens were collected. By this means, it has been possible to establish, for the first time, a precise upper limit to the epoch of their appearance in Europe. 

Nyctereutes megamastoides (Pomel) from the Villafranchian of the Auvergne and from Saint-Vallier presents cranial and dental characters sufficiently close to those of the late Pliocene canid from Perpignan (Roussillon), described by Depéret under the specific name of Canis donnezani belonging to the same genus Nyctereutes. The extinction of the European "Nyctereutes" group seems due to the too great alimentary specialization of this canid, whereas the Asiatic lineage represented in the Villafranchian by Nyctereutes sinensis Schlosser and at present by Nyctereutes procyonider Gray was able to maintain itself probably by means of a profound change in its alimentary regime. 

The study of the bovids from Al Jadidah (Hofuf Formation, Saudi Arabia) confirms that the fauna comes from a pre-Hipparion level. The Al Jadidah age is close to that of Fort Ternan (14 m.y.) and Beni Mellal, but cannot be older than that of Fort Ternan. The age of the Hofuf Formation is close to but slightly older than the oldest deposits of the Ngorora Formation (Kenya). 7 to 9 species have been recorded, of which 2 to 4 remain indeterminate. If the great specific diversity of te bovids from this locality gives evidence of immigrations from anterior Asia (Turkey) (e.g. Pachytragus Iigabuei sp. nov.), the bovid assemblage of Al Jadidah results in fact from a double influence: from the anterior Asia and mainly from Africa (e.g. the Caprotragoides lineage and the Neotragini? Homoiodorcas). The Al Jadidah bovids reflect, on the whole, the predominant character of open to very open environment, which supports the conclusions drawn from our two preliminary studies. 

The East African Miocene Macroscelidea, lnsectivora and Chiroptera are revised on the basis of new material. New taxa proposed are: Miorhynchocyon, .n. gen. (Macroscelididae): Míorhynchocyon meswae, n. sp.: Pronasílío ternanensis. n. gen.. n. sp. (Macroscelididae); Hiwegicyon juvenalis, n. gen. n. sp. (Macroscelididae); Parageogale, n. gen. (Tenrecidae): Prochrysochlorinae, n. subfam. (Chrysochloridae): Propottininae, n. subfam, (Pteropodidae); Chamtwaria pickfordi, n. gen., n. sp. (Vespertilionidae). Gymnurechnínus songhorensis is synonymised with G. camptolophus. The new material provides additional information on the dentition, especially of Myohyrax oswaldi. Galerix africanus. Amphechínus rusingensis, Protenrec tricuspis and Parageogale aletris. Partial skulls are described of Amphechinus rusingensis, Protenrec tricuspis, Prochrysochloris míocaenicus and Taphozous incognita. The oldest member of the Macroscelidinae (Pronasilio) is described from Fort Ternan. Galerix africanus is closely related to G. exilis from Europe. Amphechinus rusingenesis is compared with Asiatic Oligocene Erinaceinae. The Miocene age of Crocidura is rejected. On the evidence of humeri, the following families of Chiroptera are newly reported: Pteropodidae, Nycterididae, Vespertilionidae, Molossidae. Propotto is regarded as an offshoot from the Pteropodidae, not ancestral to modern forms. Chamtwaria is a primitive vespertilionoid, provisionally placed in the Kerivoulinae. Erinaceidae probably entered Africa at the beginning of the Miocene, before 20 Ma. Faunistic differences between deposits are largely to be ascribed to differences in local environment. 

The classifications of the recent vespertilionine bats were made wihtout taking in account the teeth morphology; this resulted in a reduction of the possibilities of comparison with the available fossils. The generalized use of dental formulae was abusive: this contributed to the admission of artificial genera. These conditions have long delayed the consideration of characters able to frame the phylogeny of the sub-family. In the first part of the study, the teeth morphologies are described and analysed. morphological reference types are established for each upper and lower tooth: they should make an easier elaboration of criteria for the differentiation at generic level. The position of the species in view of these criteria allows one to group them into homogeneous genera, and to appreciate the degree of relationship that the latter have between them. The second part of the study (next publicationà will develop inferences dealing with systematics and phylogeny 

La faune de Rongeurs d'Escamps (Lot) bien que relativement pauvre en espèces (dix) s'avère riche d'enseignement pour les Rongeurs de l'Oligocène inférieur d'Europe Occidentale. Cette periode semble caracterisée par une cladogenèse des Théridomyines. A Escamps, un nouveau genre (Patriotheridomys) est décrit ainsi qu'une nouvelle espèce de Theridomys. Avec Oltmamys platyceps, décrit ici plus complètement et désormais bien situé stratigraphiquement, les deux espèces précédentes constituent un ensemble original du Sud de la France. A la même époque, en Angleterre, Allemagne et Espagne, des espèces «régionales» de Théridomyinés se dersifient.  A côté de ces formes qui ne semblent pas franchir la «Grande Coupure» sont représentées deux lignées d'Issiodoromyinés (Elfomys sp et Pseudoltmomys cuvieri), une de Théridomyiné (T. (Blainvillimys) rotundidens) ainsi qu'un Gliridé, Gliravus priscus (que l'on différencie nettement de la deuxième lignée des Gliravus oligocènes : G.  meridionalis -> G. majori). On retrouve ces formes dans les gisements plus récents du niveau de Hoogbutsel où elles sont à peine plus évoluées. Le fait que les degrés évolutifs de ces lignées soient très proches laisse supposer que le laps de temps séparant les niveaux d'Ecamps et Hoogbutsel fut relativement court. 

The object of this work is the study of the artiodactyls of the karstic locality of Lissieu near Lyon. Among the sixteen species counted, five (Hyperdichobune nobilis, Hyperdichobune sp., Haplabunodon mulleri, Tapirulus depereti and Pseudamphimeryx schlosseri) are cited for the first time in this fauna. The description of part of the anterior dentition of «Dichobune» langi permits the referal of this species to the genus Hyperdichobune. The new selenodont species Dichodon lugdunensis  described in this work belongs, with its ancestor Dichodon simplex, to a lineage distinct from that of Dichodan cartieri and Dichodon ruetimeyeri. Finally, the taxonomic reality of the Lissieu species, Lophiobunodon rhodanicum Deperet, is justified. All the other species have been compared with those of Egerkingen. Even though with most of them no difference in age can be shown between the two deposits, some however indicate clearly by their degree of evolution that Lissieu is slightly more recent than Egerkingen. ln the history of Eocene mammals, the Lissieu fauna, because of its homogeneity, can be considered as a sure reference fauna of the Auversian period. 

Les Carnivores Fissipèdes de Sainte-Néboule appartiennent tous au genre Cynodictis et semblent constituer une population homogène. Celle-ci se distingue suffisamment des espèces déjà décrites pour constituer un taxon particulier : Cynodictis lacustris neboulensis n. s. sp. . L'étude des variations à l'intérieur de cette population nous a conduit à reconsidérer les critères utilisés pour définir les espèces existantes et à regrouper certaines d'entre elles. Il semble qu'il demeure cependant trois lignées distinctes dans le genre Cynodictis mais le matériel nous paraît encore insuffisant pour traduire cette remarque en termes de systématique. 

The recently discovered, very rich, Pliocene locality of Layna (Soria, Spain), has already yielded 30 species of mammals. Hipparion fissurae, described here is more dolichopodic than other Hipparion. It is related to Hipparion crusafonti from Villaroya (Villafranchian), but more slender between other characters.
  

Perutherium altiplanerise THALER, 1967 from the Late Cretaceous of Peru has long been recognized as South America's oldest known placental mammal. Since its description Perutherium has been generally regarded as having condylarth affinity Based on our identification  of a unique notoungulate synapomorphy we recognize Perutherium as the oldest and the most generalized known member of that order. This new determination and the large taxonomic diversity (five families) of notoungulates in rocks of Paleocene age in Argentina and Brazil, favor a South American origin for this group. The occurrence of notoungulates in rocks of Late Paleocene age in Asia and North America is explained by dispersal of a notoungulate stock from South America to North America and from there to Asia.