The Tertiary of the Salla-Luribay basin consists of red beds affected by the second period of the andine compression, of Miocene ending age. The Tertiary layers are exposed at an approximate elevation of 3.500 to 4.000 meters. Two stratigraphic units can be distinguished in them: the Luribay conglomerates, in which vertical clifts result from erosion, and the Salla layers consisting mostly of consolidated clays. These clays are very fossiliferous and have furnished a rich vertebrate fauna which gave to R. Hoffstetter the possibility to establish the Oligocene age of these beds. Sediments of same age has been reported to be present in several other places of Bolivia, particularly near Lacayani, where have been collected highly hypsodont Rodents, different from those found in Salla-Luribay basin. 

The eggshell of Nicobar scrubfowl (Megapodius nicobariensis) is described for the first time. Its egg porosity is calculated and discussed with data from several taxa (another megapode, some extant and fossil reptiles including a titanosaur group) in order to compare incubation types with eggshell structure. Eggshell microstructure reflects first phylogenetic traits and does not seem to have developed major adaptative features due to the incubation conditions, except for the pore canals. 

Description, for the first time, of the skull of Ruscinomys Depéret on the basis of a nearly complete specimen, and description of a new facial part of a Megacricetodon Fahlbusch skull (material from upper Miocene, Spain). New description of the skull (facial part) of " Cricetodon" incertum Schlosser on the basis of the specimen from the Oligocene of Quercy phosphorites already published by S. Schaub.
Comparison of the skull of “ C. ” incertum with that of the asiatic genus Cricetops Matthew and Granger and that of the North-American Eumys Leidy. These three genera of similar age display very distinct characteristics; their common origin must go back to the Eocene, as Wood believed.
Comparisons of the auditory regions of Ruscínomys and of Megacricetodon where important differences are noted. Comparison of the Cricetodontines from European Miocene with contemporary Cricetines. Contrary of the opinion held by Schaub, their anatomical differences do not seem suflicient to warrant the contention that none of the Cricetines derive from a form close to one of the known Cricetodontines. 

The review of material recently collected in the new localities from the “Phosphorites du Quercy", and different localities from the South of France, bring new informations on the genus Eucricetodon THALER. 1966, and Pseudocricetodon  THALER,  I969 (Middle and Upper Oligocene. Western Europe). Thanks to Eucricetodon huerzeleri VIANEY-LIAUD, 1972, which were unsufficiently known until now, is proposed. During the middle Oligocene Eucricetodon atavus  MISONNE, 1957 seems to give rise to two lineages. One of them led to Eucricetodon huberi,which however exhibits a larger size and a development of progressive characters on the teeth. The other would be Eucricetodon huerzeleri well differentiated at the “Mas de Pauffié" standard level (beginning of the upper Oligocene). The ornementation of lower incisors is described, when possible. Though the fossils are not abundant, it seems that the ancestral lineage, Eucriretodon atavus, remains in  the upper Oligocene (Boningen standard level). evolving into Eucricetodon praecursor SCHAUB, 1925 (Rickenbach standard level). The characters of Eucricetodon dubius  (SCHAUB. 1925), represented by a numerous population in Pech Desse and Pech du Fraysse (Quercy). confirm that this species and Eucricetodon praecursor  belong to two different lineages. As Eucricetodon dubius shows more primitive features, this species could not originale from Eucricetodon atavus -Eucricetodon huberi. The appearance of this species at the level of Mas de Pauffié could be the result of an immigration. A new definition of Pseudocricetodon incertus (SCHLOSSER. 1884) is given. This species has been found in several localities, where it had not been identified until now. lts comparison with Pseudocricetodon moguntiacus  (BAHLO. l975), found at several localities from the standard level of Antoingt (end of middle Oligocene). shows a parallel evolution to that of Pseudocetodon incertus, which is  of larger size and with a less complicated pattern of teeth. 

In this article is described a new species of Nyctitheriidae with primitive characters: Leptacodon nascimentoi n. sp. from the early Eocene of Silveirinha (Portugal). 

A strong lophodonty, an extreme heterodonty, some hypsodonty and regular overlayings of coronal cement are prominent features of the genus Leptolophus (Palaeotheriinae = Palaeotheriidae s.s.). The histological pattern of the teeth unusually joins type II enamel prisms, characteristic of advanced ungulates, together with archaic features, such as an almost complete lack of Hunter-Schreger zonation and a weak expanse of peritubular dentine. The skull is narrow and slender, with an elongated ante-orbital facial region, a moderately notched nasal aperture, a rather elongated post-canine diastem, parallel zygornatic arches and a fairly dorsally located squamoso-mandibular joint.The functional analysis brings to light "ectolophodont" masticatory cycles with two phases, in which maximum power was applied, contrary to equíds, on hindmost teeth; likewise, skull accomodations to increasing height of the teeth are quite different. This study leads to the assumption that Leptolophus may have been light mammals, living in rather open surroundings, browsing on herbaceous plants or leaves cropped close to the ground. Moreover, it appears that it could have been some inadequacy of dental structures to the dietary, which leaded to quick wear of the teeth and to many enamel notches, but had been somewhat balanced by the early increase of hypsodonty, not induced in such a case by a biotop deterioration (as it will happen at the end of the Eocene). This ínadaptation might account for the short duration of the genus Leptolophus, whose the 3 species, L. stehlini, L. nouletí and L. magnus n. sp. are indeed confined in the level MP 16. Its geographical spreading (as far as known, South of western Europe) and the morphological pattern of its dentition suggest that this genus would have been related to early upper Eocene endemic spanish forms.





  

The general principles guiding the study of wear facets which develop during mastication in mammals possessing tribosphenic molars are named. The application of this method of study to the molars of European Tertiary Didelphidae shows that the lineage of this family as represented by the species Peratherium cuvieri (Upper Eocene), P. elegans (Lower-middle Oligocene) and P. antiquum (Upper Oligocene) has propessively evolved toward a more carnivorous diet. 

An unpublished mandible of the large hyracoid Titanohyrax andrewsi from the early Oligocene Jebel Qatrani Formation, Fayum Depression, Egypt is described. This specimen has a twofold importance. Firstly, it opens an unexpected window on early paleontological research in the Fayum because it was discovered as early as 1904 by the French paleontologist René Fourtau during an expedition to the Fayum organized by the Muséum National d’Histoire Naturelle, Paris (MNHN). This expedition has remarkably never been mentioned in the literature. Secondly, the mandible documents the best-preserved specimen of T. andrewsi, permitting a revision of one of the very rare Paleogene hyracoids. Interestingly, the new mandible was discovered two years before the first report of the species by Charles W. Andrews. The hypodigm of T. andrewsi is reviewed and the dentition as a whole is compared in detail, notably with other Titanohyrax species from the Fayum. The validity of the large Titanohyrax “schlosseri” species is discussed, but a pronounced sexual size dimorphism for T. andrewsi is favoured. 

We report the first confirmed fossil bats from North Dakota, including a new species referable to the Vespertilionidae represented by a maxilla with P4-M3 from the Brule Formation, Fitterer Ranch local fauna, early Oligocene, Whitneyan North American Land Mammal Age. Unassociated postcranial fragments of the humerus and femur also represent a vespertilionoid, but appear to reflect a different, unidentified species. The new taxon, Quinetia frigidaria sp. nov., is referred to the genus Quinetia, previously known only from approximately contemporaneous deposits in Europe. The new species is larger than Quinetia misonnei from the early Oligocene of Belgium. It is similar in some morphological characters to Chadronycteris rabenae (Chiroptera incertae sedis) of the late Eocene (Chadronian) of northwestern Nebraska and to Stehlinia species (?Palaeochiropterygidae) from the Eocene and Oligocene of Europe, but differs from each in morphological details of the dentition and maxilla. An unassociated talonid of a lower molar from Fitterer Ranch shows myotodont morphology, unlike the nyctalodont lower molars in Q. misonnei, and thus represents a second chiropteran taxon in the fauna. Quinetia frigidaria is a member of a Paleogene radiation of bats near the low point of the Eocene-early Oligocene decline in global temperatures, increased seasonal aridity, and loss of tropical floras from mid-latitude North America. 

Record of life and works of Henri Menu, French zoologist, contributor to the knowledge of living and fossil bats. 

 Detailed anatomical description of arsinoithere cranial remains from the Lower Oligocene, Fayum Depression, Egypt, provides the basic data for a systematic investigation. All cranial and some postcranial features are assessed from a phylogenetic standpoint. Several soft tissue characters are then added to a cladistic analysis based on 54 derived ungulate morphological characters. The resulting phylogenetic hypothesis implies that perissodactyls, sirenians, proboscideans and arsinoitheres constitute a monophyletic unit (5 synapomorphies). However, increasing the tree length by 3 steps reveals a closer association between hyraxes and perissodactyls. Nevertheless, 13 synapomorphies link proboscideans, sirenians and  arsinoitheres to the exclusion of all other ungulates. Form of the sphenopalatine and ethmoid foramina, recurved posttympanic process, absence of a fenestra rotundum in the petrosal, vestigial paroccipital process of the exoccipital and the highly unusual absence of a hypoglossal foramen in the skull, imply a robust sister-group relationship between arsinoitheres and proboscideans. In this analysis artiodactyls share only one derived character with all other ungulates studied. Monophyly of Ungulata, including Artiodactyla, is therefore only weakly supported. It is argued that pedal anatomy of hyraxes is non-homologous with that of Tethytheria. Arsinoitherium should now be classified within Tethytheria, sharing a sister-group relationship with Proboscidea. Hyraxes are excluded, thus refuting the concept of Paenungulata. However, monophyly of the wider concept, Pantomesaxonia, containing hyraxes, perissodactyls, sirenians, proboscideans and now, arsinoitheres, is supported by this study. 

Abstract not available 

Two species lineages of North American late Paleocene Plesiadapis exhibit a pattern of size divergence from a common ancestral lineage. Time series of fossils in each of these lineages are analyzed to test the idea that size divergence represents competitive character displacement. The critical factor in a test of character divergence is showing that divergent lineages evolved directionally rather than randomly (multifactorially). Analysis of evolutionary rates and their temporal scaling in Plesiadapis shows that both divergent species lineages have the scaling slope expected for lineages evolving randomly rather than directionally, and size divergence in Plesíadapis does not represent character displacement. Rates of evolution commonly observed on a per-generation time scale are high enough to produce character displacement within a few generations. Thus character displacement is not likely to be visible on scales of time that can be studied in the fossil record. 

A new artiodactyl, Diacodexis antunesi n.sp., is described from the early Eocene of Silveirinha, Portugal. Comparisons are made with Diacodexis gazini GODINOT, 1978, D. varleti SUDRE et al., 1983, D. cf. varleti from Paris Basin sites, D. sp. from Dormaal and from localities in Spain and England, D. secans from North America and D. pakistanensis from Asia; affinities and evolutive tendencies are discussed. The presence of Diacodexis in the locality of Silveirinha confirms the very early Eocene age of the latter. As Diacodexis antunesi appears ta be more primitive than D. gazini from Rians (early Eocene of France), it lends corroboration to the interpretation (essentially based previously on condylarths) of the Silveirinha assemblage as the oldest Eocene fauna known in Europe and supports the hypothesis that early artiodactyls migrated from Europe to North America. 

The study of three new populations of Plesiosminthuspromyarion from the "phosphorites du Quercy" and of material from "Auvergne" brings new data on european oligocene Dipodidae. They appear in Western Europe at the beginning of late Oligocene. Evolutionary trends of the group are drawn and particularly the emergence of morphotypes announcing P. schaubi, from the Coderet level, is revealed among the most recent populations of P. promyarion. Differences are attempted to be drawn between the three species : P. promyarion, P. myarion and P. schaubi by restudying the type-population of P. myarion from the aquitanian deposits of Chavroches (Allier) in comparison with two other populations from the same age and the same region. Relationships between early european, american and asiatic Dipodidae are discussed. 

The teeth and the astragale of the Creodonta from Ste-Néboule (Lot) are referred to Hyaenodon brachyrhynchus. Isolated teeth fit morphologically the material from La Débruge (Middle Ludian). However, biometric analysis suggests an Upper Ludian age. 

The family Didelphidae is represented by three species in the Sainte-Néboule site, phosphorites of Quercy (lower Oligocene, San Cugat's nivel): Amphiperatherium minutum (Aymard), Amphiperatherium sp. and Peratherium cuvieri (Fischer). Only the first  and  third species are abundant (88 and 97 pieces). This two populations are described. The marsupial fauna of the european lower Oligocene is not recognized in its entirety in this site. 

The description of new material from three classic middle Eocene localities of western Europe permits the addition of details to the systematics of primitive Pseudosciurids. The cranial anatomy of Protadelomys cartieri (STEHLIN and SCHAUB) from Egerkingen is described here and compared to that of the Adelomyines from the late Eocene, until now the only ones known. The morphologic and biometric study of the dentition of P. cartieri compared to that of P. alsaticus n. sp. from Bouxwiller and P. Iugdunensis n. sp. from Lissieu, forms respectively older and younger than P. cartieri, permits the evolutionary tendencies of the group to be demonstrated and shows that notable differences in age exist between these localities. This ensemble of forms can constitute a valuable guide lineage in the establishment of a fine stratigraphy of the period. Other less well known lineages are present at Egerkingen along with P. cartieri. They can be related to genera that have been noted int he late Eocene. In conclusion, a criticism of recent zonation proposals, made by divers authors, completes this article. 

A new Tertiary megadermatid is described from Rackham's Roost Site, a Pliocene limestone cave deposit on Riversleigh Station, northwestern Queensland, Australia. It appears to represent the first Australian record of Megaderma GEOFFROY, 1810, a genus otherwise known from Tertiary African and European taxa and the living Asian species M. spasma (LINNAEUS, 1758) and M. (Lyroderma) lyra PETERS, 1872. Megademza richardsi n. sp. is one of the smallest megademiatids known. It exhibits a mixture of plesiomorphic and autapomorphic features, the latter appearing to exclude it from being ancestral to any living megadermatid. The new species is one of eight megadermatids identified from the Australian fossil record, most of which are referable to Macroderma MILLER, 1906. 

Avant propos au "Premier catalogue des specimens-types Paléontologiques déposés dans les collections de l'Université de Montpellier II".