Fourteen distinct phyletical lineages which belong at least in three families: Ischyromyidae ALSTON, 1876, Gliridae THOMAS, 1896 and Theridomyidae ALSTON, 1876, have been identified after the study of more than 3600 rodent dental remains from about twenty Early and Middle Eocene european localities. A systematical and phylogenetical revision of these rodents has been achieved. Nearly all the specific and generic diagnosis are emended. Several new combinations and synonymies are proposed. Four new species and two new genera, Euromys nov. (Ailuravinae) and Hartenbergeromys nov. (Microparamyini), are named and described. Euromys nov. gen. is known by three distinctive ypresian (MP 7 to MP 10 european reference levels) chronospecies. This new lineage is thought to be the direct ancestor of Meldimys MICHAUX, 1968 and Ailuravus RUTIMEYER, 1891. A new species of the genus Plesiarctomys BRAVARD, 1850, Pl. lapicidinarum from Condé-en-Brie (MP 8-9 reference level), allows to relate the Plesiarctomys lineage to the Pseudoparamys MICHAUX, 1964 one. The taxa Sparnacomys HARTENBERGER, 1971, Pantrogna HARTENBERGER, 1971, and Corbarimys MARANDAT, 1989 are erected to genus rank; the last one is not thought to be an Ischyromyidae. A new chronospecies of Pantrogna, P. marandati nov. sp. from the locality of Prémontré (MP 10 reference level), is described. This lineage is at the origin of two others, namely Masillamys TOBIEN, 1954, including M. mattaueri (HARTENBERGER, 1975) nov. comb. (MP 10 reference level), and Hartenbergeromys nov. gen., known from MP 10 (H. hautefeuillei nov. sp.) and MP 11 (H. parvus TOBIEN, 1954) reference levels. The phylogenetical position of Hartenbergeromys nov. gen., at the origin of the european family Theridomyidae, is discussed. The systematical and phylogenetical status of two probable Paramyinae, "Paramys" woodi MICHAUX, 1964 and an unnamed genus and species, are discussed. New populations of the primitive Gliridae Eogliravus HARTENBERGER, 1971 and of the primitive Theridomyidae Protadelomys HARTENBERGER, 1968, are described and assigned to previously known species.

  

On the Track of Ice Age Mammals justifie pleinement son titre car l'auteur, Anthony J. Sutcliffe, apporte au lecteur faits et interprétations qui l'amèneront à s'intéresser encore plus au passé récent et à l'avenir de son environnement et à la question de l'impact de l'homme sur la nature. Après les chapitres qui présentent les temps glaciaires et les divers témoignages qui nous en sont parvenus, les cinquième et sixième apportent les informations nécessaires à la compréhension des résultats que nous donnent les chercheurs: principes, moyens d'étude et limites des méthodes, difficultés de l'intégration des données dans un cadre stratigraphique, variabilité des signaux climatiques, variabilité de leur intensité selon l'endroit par rapport au centre de la glaciation, complications liées à la qualité inégale de l'enregistrement géologique, en mer et sur le continent. 

A systematic analysis of an hipparionine horse assemblage from Maragheh, Iran is made. A brief orientation to systematic philosophy and informal superspecific characterizations of some Old World hipparionines is given as a background to this work. A character state analysis of skulls is made, and has revealed five distinct species. A character state and stratigraphic trend analysis of isolated check tooth and postcranial remains, with known provenance, is also made. These two combined analyses reveal that the most resolute discrimination of hipparionine species and their evolutionary relationships occurs when multiple character complexes of associated skulls, maxillary and mandibular dentitions are made. When this is not possible, skulls have provided the best basis for discriminating species and their evolutionary relationships. Traditional characters of isolated cheek teeth and postcranial remains are shown here to offer limited information content for hipparionine phylogenetic systematics. The systematic portion of this study includes a comprehensive description of cranial and postcranial remains, and has further corroborated the distinction of five species which belong to at least three superspecific groups including: «Hipparion» geltyi sp. nov., Group 1; Hipparion prostylum (s. l.), and Hipparion campbelli sp. nov., Group 3; «Hipparíon» aff. moldavicum and «Hipparion» ?matthewi, Group 2. These species stratigraphic ranges and evolutionary relationships are also given here and argued to be important for establishing future hipparionine geochronologic correlations between a number of Eurasian late Miocene provinces. 

A new example of parallelism in the dental pattern ofthe Theridomyidae is illustrated by the description ofa new species: Theridomys Iudensis from the standard level of Antoingt (middle Oligocene). Considering the occurence ofthis parallelism phenomenon. the use of numerous qualitative and quantitative criteria is essential to characterize the different stages ofthe different evolutive lineages. Thus, a new simple parameter is proposed (CHY = H+l/0,5 L) to estimate hypsodonty of the medium hypsodont Rodentia. 

The genus Parasteiromys AMEGHINO, 1904 is revalidated, and P. friantae sp. nov. (Hystricognathi, Erethizontidae) from Colhuehuapian (early Miocene) sediments of the southern cliff of Colhue-Huapi Lake (Province of Chubut, Argentina), is described. The molar morphology of these taxa and of living porcupines adds new elements to understand the dental evolution of the Erethizontidae, and to propose the hypothetical ancestral molar pattern for this family. This pattern does not correspond to any of the morphologies traditionally proposed as ancestral for South American hystricognathous rodents. The proposed pattern is characterized by a metaloph disconnected from the posteroloph and oriented towards the hypocone, and the third loph incompletely developed with the lingual portion homologous to the mesolophule of Baluchimyinae (Chapattimyidae) from the Miocene of Pakistan. The inferred steps of the molar evolution of erethizontids towards the pentalophodont condition, considered derived for the family, are illustrated. This study strengthens the hypothesis placing erethizontids in a basal position among rodents of the suborder Hystricognathi.

  

 Detailed anatomical description of arsinoithere cranial remains from the Lower Oligocene, Fayum Depression, Egypt, provides the basic data for a systematic investigation. All cranial and some postcranial features are assessed from a phylogenetic standpoint. Several soft tissue characters are then added to a cladistic analysis based on 54 derived ungulate morphological characters. The resulting phylogenetic hypothesis implies that perissodactyls, sirenians, proboscideans and arsinoitheres constitute a monophyletic unit (5 synapomorphies). However, increasing the tree length by 3 steps reveals a closer association between hyraxes and perissodactyls. Nevertheless, 13 synapomorphies link proboscideans, sirenians and  arsinoitheres to the exclusion of all other ungulates. Form of the sphenopalatine and ethmoid foramina, recurved posttympanic process, absence of a fenestra rotundum in the petrosal, vestigial paroccipital process of the exoccipital and the highly unusual absence of a hypoglossal foramen in the skull, imply a robust sister-group relationship between arsinoitheres and proboscideans. In this analysis artiodactyls share only one derived character with all other ungulates studied. Monophyly of Ungulata, including Artiodactyla, is therefore only weakly supported. It is argued that pedal anatomy of hyraxes is non-homologous with that of Tethytheria. Arsinoitherium should now be classified within Tethytheria, sharing a sister-group relationship with Proboscidea. Hyraxes are excluded, thus refuting the concept of Paenungulata. However, monophyly of the wider concept, Pantomesaxonia, containing hyraxes, perissodactyls, sirenians, proboscideans and now, arsinoitheres, is supported by this study. 

Cranial anatomy of a late Miocene rhizomyid, Brafhyrhizomys cf. B. pilgrimi, provides new evidence on the origin of the dorsal, round infraorbital foramen of living rhizomyines. Primitive rhizomyids retain a myomorphous keyhole foramen with a long ventral slit that retracts upward during the evolutionary history of the Rhizomyidae. The primitive condition of the elongated ventral slit is represented by Kanisamys sivalensis. Among later burrowers the foramen shows progressive dorsal migration, the ventral slit terminating midway up the snout in B.tertracharax and B. choristos ; well above the midline of the snout in Brachyrhizamys cf. B. pilgrimi. Apparently this transformation began earlier among Rhizomyinae than among Tachyoryctinae and continued to a more derived stage in rhizomyines. ln living Rhizomyx the ventral slit is absent and only a high round hole remains at the anterior end of the zygomatic arch. 

The classifications of the recent vespertilionine bats were made wihtout taking in account the teeth morphology; this resulted in a reduction of the possibilities of comparison with the available fossils. The generalized use of dental formulae was abusive: this contributed to the admission of artificial genera. These conditions have long delayed the consideration of characters able to frame the phylogeny of the sub-family. In the first part of the study, the teeth morphologies are described and analysed. morphological reference types are established for each upper and lower tooth: they should make an easier elaboration of criteria for the differentiation at generic level. The position of the species in view of these criteria allows one to group them into homogeneous genera, and to appreciate the degree of relationship that the latter have between them. The second part of the study (next publicationà will develop inferences dealing with systematics and phylogeny 

The review of material recently collected in the new localities from the “Phosphorites du Quercy", and different localities from the South of France, bring new informations on the genus Eucricetodon THALER. 1966, and Pseudocricetodon  THALER,  I969 (Middle and Upper Oligocene. Western Europe). Thanks to Eucricetodon huerzeleri VIANEY-LIAUD, 1972, which were unsufficiently known until now, is proposed. During the middle Oligocene Eucricetodon atavus  MISONNE, 1957 seems to give rise to two lineages. One of them led to Eucricetodon huberi,which however exhibits a larger size and a development of progressive characters on the teeth. The other would be Eucricetodon huerzeleri well differentiated at the “Mas de Pauffié" standard level (beginning of the upper Oligocene). The ornementation of lower incisors is described, when possible. Though the fossils are not abundant, it seems that the ancestral lineage, Eucriretodon atavus, remains in  the upper Oligocene (Boningen standard level). evolving into Eucricetodon praecursor SCHAUB, 1925 (Rickenbach standard level). The characters of Eucricetodon dubius  (SCHAUB. 1925), represented by a numerous population in Pech Desse and Pech du Fraysse (Quercy). confirm that this species and Eucricetodon praecursor  belong to two different lineages. As Eucricetodon dubius shows more primitive features, this species could not originale from Eucricetodon atavus -Eucricetodon huberi. The appearance of this species at the level of Mas de Pauffié could be the result of an immigration. A new definition of Pseudocricetodon incertus (SCHLOSSER. 1884) is given. This species has been found in several localities, where it had not been identified until now. lts comparison with Pseudocricetodon moguntiacus  (BAHLO. l975), found at several localities from the standard level of Antoingt (end of middle Oligocene). shows a parallel evolution to that of Pseudocetodon incertus, which is  of larger size and with a less complicated pattern of teeth. 

Le volume 11 du «Handbuch der Paläeoherpetologie», consacré aux «Serpellles» et redigé par Jean-Claude Rage, apporte une remarquable information sur ce groupe particulier de reptiles dont on ne pouvait guere se faire une idée très précise à partir, par exemple, des seuls chapitres des traités classiques de Paléontologie et de Zoologie. 

The genus Cryptomeryx SCHLOSSER, 1886, inusited during a long period, has been discovered in Lower and Middle Oligocene localities of the Quercy region (South-West France). This new material, as well as specimens from the old collections referred to Cryptomeryx, are described; their study, allows us precising the definition of the genus, and confirming its allocation to the Tragulidae family. The type species of the genus, Crypmmeryx gaudryi (= Lophiomeryx gaudryi FILHOL, 1877), occurs in several localities at the base of the Middle Oligocene (Itardies, La Plante 2, Roqueprune 2, Soulce, Herrlingen 1). The new species C. matsoui n. sp. has been defined in the older locality of Mas de Got (top of Lower Oligocene). It is possible that the species Pseudamphimeryx decedens STEHLIN, 1910 pertains to the same genus. Also to the Tragulids must be referred the monospecific genus Iberomeryx (I. parvus GABOUNIA, 1964) from Upper Oligocene of Benara (Georgie, URSS), with which Cryptomeryx is related. These genera are not direct ancestors of Miocene tragulids; their occurrence in the Western European Oligocene results from a first immigration wawe of the family. These Tragulids are one of the most archaic groups of Ruminants. They are probably derived from a primitive stock which had acquired in Asia the selenodont condition of the dentition. 

The study of the bovids from Al Jadidah (Hofuf Formation, Saudi Arabia) confirms that the fauna comes from a pre-Hipparion level. The Al Jadidah age is close to that of Fort Ternan (14 m.y.) and Beni Mellal, but cannot be older than that of Fort Ternan. The age of the Hofuf Formation is close to but slightly older than the oldest deposits of the Ngorora Formation (Kenya). 7 to 9 species have been recorded, of which 2 to 4 remain indeterminate. If the great specific diversity of te bovids from this locality gives evidence of immigrations from anterior Asia (Turkey) (e.g. Pachytragus Iigabuei sp. nov.), the bovid assemblage of Al Jadidah results in fact from a double influence: from the anterior Asia and mainly from Africa (e.g. the Caprotragoides lineage and the Neotragini? Homoiodorcas). The Al Jadidah bovids reflect, on the whole, the predominant character of open to very open environment, which supports the conclusions drawn from our two preliminary studies. 

Perutherium altiplanerise THALER, 1967 from the Late Cretaceous of Peru has long been recognized as South America's oldest known placental mammal. Since its description Perutherium has been generally regarded as having condylarth affinity Based on our identification  of a unique notoungulate synapomorphy we recognize Perutherium as the oldest and the most generalized known member of that order. This new determination and the large taxonomic diversity (five families) of notoungulates in rocks of Paleocene age in Argentina and Brazil, favor a South American origin for this group. The occurrence of notoungulates in rocks of Late Paleocene age in Asia and North America is explained by dispersal of a notoungulate stock from South America to North America and from there to Asia. 

The revision of the old collections of fossil birds from the “Phosphorites du Quercy” and the study of new material give the following results (Gruiformes, Cariamae) :  The humeri and most of the carpometacarpi described under the name Filholornis belong in Elaphrocnemus. The ulnae ascribed to Fïlholornis belong in Idiornis. Most of the post-cranial elements of the genera Elaphrocnemus and Idiornis are described and show great similarities with recent Cariamidae and Opisthocomidae, and fossil Bathornithinae.  A new genus and a new species, Oblitavis insolitus, are created in the sub-family Idiornithinae; two new species are described in the genera Elaphrocnemus (E. brodkorbz) and Idiornis (I. itardiensis), and the species Elaphrocnemus gracilis is transferred to the genus Idiornis.  The genus Propelargus Lydekker is transferred from the family Ciconiidae to Cariamidae.  A new generic name, Occitaniavis, is created for the species Geranopsis elatus, which belong in Cariamidae, while the type-species of the genus, Geranopsis hastingsiae, is a member of the Gruidae.  The affinities between the Quercy avifauna and the Neotropical one is emphasized by the occurrence of Phorusrhacidae, previously known only from the Cenozoic of South America and the Late Pliocene or Early Pleistocene of North America. Thanks to the material collected during the new excavations, the stratigraphical position of most of the species is stated precisely, and evolutionary lineages are outlined. This study shows that the suborder Cariamae, presently restricted to two South American genera, was already extremely diversified during the Eocene, and widespread in Europe and North America. 

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

A reconstruction of a multituberculate endocranial cast is made on the basis of a complete natural cast prepared from the skull of Chulsanbaatar vulgaris, and other less complete endocasts, all from the Late Cretaceous of Mongolia. The multituberculate endocast is of mammalian pattern but it has retained a therapsid-like lateral profile with a deep rhombencephalon and a shallow telencephalon. It is characterized by: a heart-shaped cross-section of the telencephalon; an extensive lissencephalic neocortex; a very prominent pons placed far anteriorly; a lack of cerebellar hemispheres, and very large paraflocculi. Its structure, very different from the brains of other mammals, suggests thats the Multituberculata branched very early from the main mammalian stock. This supports Simpson's (1945) idea that the Multituberculata should be placed in a subclass of their own: Allotheria MARSH. The endocast and braincase structure show that the Multituberculata had strongly developed senses of smell and hearing. The encephalization quotient of approx. 0.55 evaluated for Ch. vulgaris is relatively high for a Mesozoic mammal. 

The Condylarths from Hainin (Hainault, Belgium) show no affinity at the generic level to those known in other Paleocene localities of Europe and North America ; they are described as new forms : Monshyus praevius n. gen., n. sp. and Prolatidens waudruae n. gen., n. sp. Monshyus praevius, discovered in only one of the levels in the excavation at Hainin, is similar to the genera Microhyus TEILHARD and Louisina RUSSELL ; with them it is included in the subfamily Louisininae (Hyopsodontidae). With respect to Microhyus and Louisina, Monshyus is distinguished by the precociously modern aspect of its upper molars, the only teeth that are referable. Prolatidens waudruae, known only by lower molars, was found in several levels in the pit at Hainin. It is an arctocyonid presenting possible relationships to the North American form Oxyprimus galadrielae ; it therefore has been provisionally attributed to the subfamily Oxyclaeninae. If this attribution is confirmed, this species will constitute the first and only representative of the group in Europe. 

The discovery of eggshells of ornithoid type is signaled for the first time in the continental Campanian and Maastrichtían of Provence. The taxonomie relationship of these eggshells and their interest in the biostratigraphical study of Upper Cretaceous of Provence are discussed. 

The Montredon local fauna yielded very rare bats, represented by damaged isolated teeth. Only a few documents are available for this period of the European Neogene. ln this poor state of knowledge, the material represents three undetermined species, a supposed molossid and two vespertilionids. 

This paper presents a preliminary list of insectivores from the Vallesian beds at Montredon (France). The associated rodent fauna has established a Vallesian age for the fauna. Eleven species belonging to the Soricidae, Talpidae, Erinaceidae, and Dimylidae are identified of which four only are referred with certainty to forms already named.