The Palaeobatrachidae until now have not been reported from France. The present note makes known the discovery of very significant remains in three Tertiary localities of France: Cernay (late Paleocene), Laugnac (late Aquitanian) and Sansan (middle Helvetian). The osteologic characters, as well as the geographic and stratigraphic distribution of the family are re-stated with added details. The Palaeobatrachidae are a European family known from the late Jurassic of Catalonia (publication in course by J. Seiffert) to the Plio-Pleistocene (pre-Mindel) of Poland and Rumania. By their way of life and certain osteologic characters they are similar to the Pipidae, but they are distinguished by profound differences. It would be rash to include them in the Pipoidea or even in the Aglossa. They are apparently an independant group, essentially holarctic, which held in Europe a role equivalent to that played by the Pipidae south of Tethys. 

This paper is about the oldest African rodents faunas, from the late Early Eocene, or early Middle Eocene, Glib Zegdou (Algeria) and Chambi (Tunisia) localities. Five species are described and figured, belonging to a new family here created, the Zegdoumyidae.

This family is compared to the Early and Middle Eocene rodents families from Asia, Europe and North America (Chapattimyidae, Yuomyidae, Gliridae, Theridomyidae, lschyromyidae and Sciuravidae), as well as to those known from the Late Eocene African locality Bir El Ater (Anomaluridae and Phiomyidae).

On the one hand, it seems clear that the African endemic Anomaluridae arise from the Zegdoumyidae. On the other hand, the lschyromyidae, or primitive Sciuravidae, may be the most reliable ancestral groups for the Zegdoumyidae. Thus, this new family can be considered as the sister group for the American Sciuravidae on the one hand, and for the European Gliridae on the other hand.

The biogeographical consequences of these phylogenetic hypotheses are discussed. A new phase of communication between Europe and North Africa is inferred, during the Early Eocene. It has been followed by a short period of endemism, allowing the adaptive radiation for the Zegdoumyidae, preceding the immigration of the Phiomyidae, during the Late Eocene, probably from Asian relatives. 

Is hereby described an impressive lchnoiauna belonging to the Lower to Middle Ludian of the Gard (S. France). The slab, already cleaned over a length of 18 m, is located near the top of the Potamides aporoschema lacustrine limestone (Lower Ludian, Euzet zone). It is therefore older than the Célas sandstone deposit, and still more than the Melanoides albigensis and M. acutus marly limestone corresponding to the Upper Levels of the Ludian stage. Although biostratigraphically older than the La Débruge and Montmartre zone, the biotope shows already a sampling of very tall Artiodactyles, Perissodactyles and Carnivorous. One of the most « majestic ›› Artiodactyles, Anopolotheriipus lavocati, nov., points out a huge size type. To mention also among the Ichnotypes described, 10, the big Perissodactyle Palaeotheriipus similimedius, nov., and the big Carnivorous Hyaenodontipus praedator, nov. 

Extensive collecting in previously known and new Middle Eocene rodent localities in the Subathu Group of the Rajauri and Reasi districts, Jammu and Kashmir (northwest Himalaya, India) has yielded over 500 isolated cheek teeth, numerous incisors and cheek tooth fragments, and two mandibular fragments with M/1-M/3. An analysis of this additional material and a restudy of that reported earlier from the Metka (Rajauri) area has revealed the presence of diverse Middle Eocene ctenodactyloid rodents in India. The assemblage comprises six genera and twelve species, viz.., Bírbalomys woodi, B. ibrahimshahi, B. sondaari, Basalomys vandermeuleni, B. ijlsti, B. lavocati, Chapattimys wilsoni, C. debruijni, Gumbatomys asifi, cf. Advenimus bohlini, cf. Petrokoslovia sp. indet. 1 and cf. Petrokoslovia sp. indet. 2. Most species of this assemblage are endemic to the Indian subcontinent and are referred to the Family Chapattimyidae. However, at least three taxa, viz.., cf. Advenimus bohlini, cf. Petrokoslovia sp.indet. 1 and cf. Pezrokoslovia sp. indet. 2 show close affinities with their Central and Eastern Asiatic contemporaries and are attributed to the Family Yuomyidae. The dentitions of most of the Middle Eocene rodents from India are described here in better detail thus providing more precise characterization than was available earlier. The taxonomie status of Basalomys, formerly a subgenus of Birbalomys has been raised to that of a genus. Basalomys vandermeuleni is here designated as the type species of Basalomys in place of B. ijlsti. The rodent assemblage from the Subathu Group is specifically similar to that from the coeval Kuldana Formation of Pakistan. The absence of Birbalomys ibrahimshahi and cf. Advenimus bohlini in Pakistan is related to sampling bias. The identifications of the Early Eocene rodents from Barbara Banda, Pakistan have been reexamined and the familial assignments of some of the Early and Middle Eocene rodents from Central and Eastern Asia have been discussed and reviewed in the light of recent discovery of early rodents in Mongolia.

  

A fragmentary mandible and maxilla of a small sized Agriotherium of a young individual discovered from a Pliocene fissure filling in Xiaoxian county (Anhuei Province, China) are described. Judging from the morphology of the dentition and its dimensions the new material can be identified as Agriotherium inlermedium (STACH l957). Hendey's proposition (1980) that the Agriotherium species are derived from Indarctos is reconsídered on the basis of the new documents. As a result of a more general phylogenetic discussion it can be stated, that: 1. the supposed size increase as well as other trends, leading from Indarctos to Agriotherium are untenable ; 2. there are no positive indications to assume a phylogenetic transition of these two genera. 3. there are no real arguments in favor of an adaptational reversal in the evolution of Agriotherium. Hence, many features of that genus supposed by Hendey to be derived are plesiomorphic ; 4. regardless of the previous points it is methodologícally impossible to establish direct ancestor - descendant relationships between Indarctos and Agriotherium species, as Hendey did. Based on the data available and especially on the characters of the new material from China it is more likely that Agriotherium and Indarctos are two genera which developed independently. While advanced Agriotherium species, e.g. A. africanum, resemble in some respects Indarctos by adaptational analogies, more primitive species, e.g. Agriotherium intermedium, are quite dissimilar to lndarctos. While Indarctos might be derived from an Ursavus like forerunner, Agriotherium has its roots more likely somewhere in between Ursavus and the Hemicyon-group. 

A comparative bone histological study of the three known genera of the endemic European ornithopod dinosaur family, Rhabdodontidae, is presented here in an ontogenetic context. Investigated specimens were assigned to different ontogenetic stages based exclusively on the histological indicators of osteologic maturation during diametrical bone growth; an entirely size-independent method as opposed to most previous studies. Qualitative comparison of bone histology of corresponding ontogenetic stages and elements among the three valid rhabdodontid genera, Mochlodon, Zalmoxes, and Rhabdodon, revealed some consistent patterns. Genus specific histological differences within Rhabdodontidae are most expressed between Rhabdodon and the Mochlodon-Zalmoxes clade. These indicate a prolonged phase of fast growth and a less constrained cyclicity in the growth dynamics of Rhabdodon, as opposed to the slower and more regulated growth strategy reflected in the bones of Mochlodon and Zalmoxes. These genus specific differences are consistent with the phylogenetic interrelation of the genera and are most probably related to the pronounced differences in body size. However, when compared to other ornithopods, most detected histological features in rhabdodontids do not seem to reliably reflect either phylogenetic relations or body size. A notable common feature of all rhabdodontid genera irrespective of body size is the ontogenetically early onset of cyclical growth and secondary remodelling; a pattern that more resembles the condition found in derived ornithopods than that described in more basal taxa which are closer relatives of rhabdodontids. The recognition of taxon-specific histological patterns as well as patterns indicative of ecological and thereby functional traits clearly requires more accurate, preferably quantitative evaluations.   

Deux types de remaniements post mortem me paraissent caractéristiques de l'état de conservation des dents de mammifères fossiles dans les Phosphorites du Quercy :

1) Des destructions localisées d'origine biologique, sous forme de galeries de morphologie très variable creusées dans la dentine et le cément, et impliquant sans doute la participation de différents types de micro-organismes. Ces altérations se sont développées peu de temps après la mort, avant la fossilísation proprement dite et se sont rapidement arrêtées après l'enfouissement dans le sédiment phosphaté.

2) Des perturbations dans les structures de la dentine liées aux variations locales de minéralisation, provoquées par une imprégnation diffuse des zones les moins calcifiées par divers minéraux et probablement surtout de l`apatite. 

The study of endocasts, or casts of the endocranial space, have played an important role in shaping understanding of mammalian, and particularly primate, brain evolution. Recently, the reconstructions of three-dimensional virtual endocasts from high-resolution computed tomography images have allowed for the visualization and quantification of endocasts in several Paleocene and Eocene primate species. Here we present the virtual endocast of MaPhQ 289 (informally known as the Montauban 9 skull), a specimen of Necrolemur antiquus Filhol 1873, a middle to late Eocene European primate of the family Microchoeridae. The virtual endocast of MaPhQ 289 reveals a lissencephalic surface morphology with expanded temporal poles and minimal overlap of the cerebellum or olfactory bulb by the cerebrum, which closely resembles the morphology of the endocast of its contemporary relative, Microchoerus erinaceus (Primates, Microchoeridae). MaPhQ 289 yields an endocranial volume (ECV) of 2.36 cm3, about 60% smaller than the volume of the most commonly cited ECV of N. antiquus. Thus, the size of the brain of N. antiquus relative to its body size is likely to be smaller than has been reported in previous literature, highlighting the importance of corroborating older ECV estimates with new evidence using 3-D imaging techniques. 

The skull remains referred to the genus Palaeotherium are recorded and described. Biometrical tests are made to elucidate intrageneric allometric relationships and to allow comparisons with various other perissodactyls. Apart from the well known shortness of post canine diastems and deepness of the narial opening, the genus is characterized by a great lengthening of the splanchnocranium, owing to a spreading of the post-orbital facial region, by a reduced area of the eye-socket and by the prevalence of the temporal muscle with regard to the masseter; this original shape of the masticatory apparatus needs to be related to the morphology of the jugal teeth and particularly to their  asymmetrical semi-hypsodonty.
    These animals, whose running ability was evidently poor, appear to have been adapted to rather closed environments, feeding on relatively soft vegetable matter; olfactory sense was likely to play a leading part in interindividual and environmental relationships. Such evolutionary trends might explain the disappearance of most of them, as clirnatic conditions deteriorated at the end of the Eocene, before the "Grande Coupure" which affected mammalian faunas at that time.
      Although the present paper is not directly concemed with phylogenetics, it invalidates the supposed ancestor-descendant relationship between P. castrense and P. magnum, and it suggests a possible emergence of the P. medium lineage from a P. siderolithicum stock. Moreover, the structure of the post-orbital facial area allows the  erection of a new sub-genus, Franzenitherium, for the  species lautricense and duvali.

  

Abstract not available 

A detailed comparative study of a complete skeleton of Bachitherium and a cladistic analysis of the sub-order Neoselenodontia lead us to propose a cladogram and a new classification of this group. The Tylopoda are the sister-group of the Ruminantia, which are chiefly defined by the fusion of the cuboid and navicular. Within this infra-order, Amphimeryx is the sister genus of a tetraselenodont group, in which the Hypertragulidae are well-separated group from a monophyletic group defined by the loss of trapezium, fusion of capitatum and trapezoid, and the isolation of the hypoconid on lower molars. The most primitive genera of this group, Lophiomeryx and Iberomeryx still have an open trigonid on the lower molars, but this is lingually closed in Archaeomeryx, sister-genus of the higher Ruminantia which have fused metatarsals and more evolved milk teeth. We divide them into two pan/orders : Tragulina (including the recent and miocene Tragulidae, and the North-American Leptomerycidae), and Pecora, with reduced lateral metacarpals and a new crest (telocristid) on the lower premolars. Within the Pecora, the upper molars of Gelocus are more primitive than those of Bachitherium (a genus with many autapomorphies in the dentition) itself more primitive than the group Prodremotherium + Eupecora, with fused metacarpals. We consider the Eupecora (including several genera without frontal appendages) to be monophyletic. 

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

The fossil mammals collected from the Eocene of Hammada du Dra (northwest Sahara. Algeria) and two fragmentary teeth from the Lutetian of M'Bodione Dadere (Senegal) are described.
The fossils from the northwest Sahara come from a lacustrian deposit dated by charophytes (Raskyella aff. pecki, Raskyella n. sp.. Maedleriella lavocati, Maedleriella sp. et ? Peckichara sp.) as Middle Eocene or perhaps Lower Eocene (Gevin, Feist and Mongereau, 1974). Several hyracoids (3 or 4) identified from this formation extends the age of the family Pliohyracidae Osborn in Africa. Three forms appear to belong in the genera Megalohyrax, Titanohyrax and perhaps Bunohyrax which have been know until now only from the lower Oligocene of the Fayum (M. gevini n. sp. ; T. mongereaui n. sp.. ? Bunohyrax or Megalohyrax indet.). Another hyracoidof small size is referred to a new genus, Microhyrax (M. lavocati n. sp.).
Helioseus insolitus n. g. n. sp. is described without ordinal assignment. Azibius (Sudre, 1975) which has been the subject of questions and interpretations is reviewed.
Only one tooth from the Lutetian of M'Bodione Dadere is complete enough to interpret. lt probably belongs to a condylarth and demonstrates for the first time, the presence of the order in Africa. The second tooth is too fragmentary for comment.
In conclusion., the paleobiogeographic role of Africa at the end of the cretaceous and the beginning of the Cenozoic is discussed. 

We present here the preliminary results from a new nesting site "Sextius-Mirabeau", discovered close to the historical centre of Aix-en-Provence in Upper Cretaceous deposits. This extensive megaloolithid nesting ground, the first large scale excavation (3225 m²) in France, yielded more than 530 eggs, some organized in clusters. Their study reveals the reproductive strategy of the megaloolithid egglayer group which indicates nesting fidelity and gregarious behaviour.  

The mammalian fauna (represented only by isolated teeth) from the early Cretaceous of Morocco includes a number of non-'therian' specimens, characterized by the antero-posterior alignement of the main molar cusps. This assemblage is very heterogenous, including forms (among them Gobiconodon palaios sp. nov.) that can be related to Laurasian triconodonts and several forms so far known only in this part of Gondwana (among them Kryptotherium polysphenos gen. et sp. nov.). The latter, in particular, exemplifies the diversity of the mammalian dental morphology in the Mesozoic, often largely underestimated. In the last part of this paper, the 'therian' affinity of the 'amphilestids' is discussed, on the basis of the arrangement of molar cusps, the interlocking mechanism and the occlusal pattem. 

Le matériel provenant de nouvelles fouilles dans les phosphorites du Quercy, soumis aux divers spécialistes, a conduit à la soutenance de plusieurs thèses d'état: sur les rongeurs (Hartenberger, Vianey-Liaud), les Chiroptères et Insectivores (Sigé), les Artiodactyles (Sudre), les Squamates (Rage) et, en partie, les Chéloniens (De Broin). Chacun dans son domaine, à côté des conclusions d'ordre évolutif, paléogéographique ou paléoécologique, a pu établir des successions stratigraphiques des gisements du Quercy qui se sont révélées largement concordantes. Ainsi la succession des faunes du Quercy est actuellement l'une des mieux établies. Dans ce contexte, les différents spécialistes ont décidé de conjuguer leurs efforts dans l'étude monographique de plusieurs gisements repérés le long de cette échelle, afin de rassembler l'information paléontologique sur des faunes bien précises et de tenter d'obtenir des indications d'ordre taphonomique. 

The two species of Sandgrouse from Quercy, Pterocles validus MILNE-EDWARDS and P. larvatus MILNE-EDWARDS, are ascribed to the genus Archaeoganga MOURER-CI-IAUVIRÉ which includes a third, very large species, A. pinguis. The sandgrouse of Saint-Gérand-le-Puy, Pterocles sepultus MILNE-EDWARDS, is ascribed to a new genus, Lepzoganga. This form appears in the Upper Oligocene of Quercy, in Pech Desse and Pech du Fraysse localities, and is still present in the Lower Miocene of Saint-Gérand-le-Puy and Paulhiac. Recent sandgrouse live in semidesert or desert areas. The indications provided by mammal and bird faunas in the localities where sandgrouse were found, confirm that the paleoenvironment was open and arid. The morphological study of these fossils indicates that, in the Upper Eocene, the Pteroclidae were already completely individualized with respect to Charadriiformes.
  

The family Didelphidae is represented by three species in the Sainte-Néboule site, phosphorites of Quercy (lower Oligocene, San Cugat's nivel): Amphiperatherium minutum (Aymard), Amphiperatherium sp. and Peratherium cuvieri (Fischer). Only the first  and  third species are abundant (88 and 97 pieces). This two populations are described. The marsupial fauna of the european lower Oligocene is not recognized in its entirety in this site. 

Bats are among the most numerous and widespread mammals today, but their fossil record is comparatively meagre and their early evolution poorly understood. Here we describe a new fossil bat from dental remains recovered from late Early Eocene sediments at Prémontré, northern France. This 50 million-year-old bat exhibits a mosaic of plesiomorphic and apomorphic dental features, including the presence of three lower premolars, a single-rooted p3, short p4 with metaconid, myotodont lower molars and a tall coronoid process of the dentary. This combination of features suggests it is an early member of Vespertilionidae, today’s most speciose and geographically widespread bat family. The Prémontré bat has bearing on hypotheses about the origins of vesper or evening bats (Family Vespertilionidae), as well as crown-group chiropterans.


  

HGL50 is a latest Early-early Middle Eocene vertebrate-bearing locality located in Western Algeria. It has produced the richest and most diverse fauna of amphibians and squamate reptiles reported from the Palaeogene of Africa. Moreover, it is one of the rare faunas including amphibians and squamates known from the period of isolation of Africa. The assemblage comprises 17 to 20 taxa (one gymnophionan, one probable caudate, three to six anurans, seven ‘lizards’, and five snakes). Two new taxa were recovered: the anuran Rocekophryne ornata gen. et sp. nov. and the snake Afrotortrix draaensis gen. et sp. nov. The locality has also yielded the first confirmed anilioid snake, the first Palaeogene gymnophionan, and probably the first caudate from the Palaeogene (and possibly from the Tertiary) of Africa. The presence of a caudate at that time in Africa would be of particular interest; unfortunately, the available material does not permit a definitive identification. The fauna comprises Gondwanan and more specifically West Gondwanan vicariants, probably autochthonous groups and a Eurasian immigrant (assuming that the identification of the caudate is accurate). The fauna from HGL50 is clearly distinguished from the few other Eocene assemblages of Africa. However, if this results largely from differences in geological ages, geographic positions of the localities and mainly differences in environments took a part in the composition of the faunas.