A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian. 

Based on material from 30 localities, morphologic dental, cranial and biometric analyses have permitted the characterization of two parallel Issiodoromyine lineages, and also the definition of diverse species representing several evolutive stages. Thus it is that new lineages complete the contribution made by the Theridomyinae and Cricetidae and permit, for the Quercy in particular, additional precision in the biochronologic succession of the localities. One of the lineages is limited to the genus Pseudoltinomys LAVOCAT; the other evolves from the genus Elfomys HARTENBERGER to the genus Issiodoromys BRAVARD in GERVAIS. The latter is affected by profound anatomical changes due to a functional modification of the mastication apparatus. These changes seem to be able to be put in relation with the aridification and cooling of the climate at the end of the Eocene. At the end of the middle Oligocene, a new chewing structure is achieved. It is found in diverse living rodents that inhabit a rather arid steppe environment (Cavia, Pedetes, Ctenodactylus). To these supposed nearby ecologic conditions, these rodents have responded in a convergent fashion. It is possible to attribute to the extreme specialization of Issiodoromys its incapacity to adapt to the new climatic crisis of the end of the Oligocene. The arrival of immigrants may be considered as another cause of its disappearance at this time, complementary or not with the first. 

Seventeen species of cricetid rodent are recognized and described from lower and middle Siwalik deposits in the Potwar Plateau of Pakistan. These species are grouped in three categories, characterized as subfamilies (e. g., Megacricetodontinae, Myocricetodontinae, and Dendromurinae); an additional and more abundant category of rodents from these deposits, the Democricetodontinae, is excluded from this study, and will be described in a later study. Fifteen of the species are new, and four new genera are described. The Siwalik cricetid taxa are : Megacricetodon aquilari, n. sp.; Megacricetodon sivalensis, n. sp.; Megacricetodon daamsi, n. sp.; Megacricetodon mythikos, n. sp.; Punjabemys downsi, n. gen. & n. sp.; Punjabemys leptos, n. gen. & n. sp.; Punjabemys mikros, n. gen. & n. sp.; Myocricetodon sivalensis, n. sp.;  Myocricetodon sp.; Dakkamyoides lavocati, n. gen. & n. sp.; Dakkamyoides perplexus, n. gen. & n. sp.; Dakkamys asiaticus, n. sp.; Dakkamys barryi, n. sp.; Dakkamys sp.; Paradakkamys chinjiensis, n. gen. & n. sp.; Potwarmus primitivus, n. gen.; and Potwarmus minimus, n. gen. & n. sp. This diverse record of middle Miocene small mammals illuminates a profound radiation of cricetid rodents in southem Asia, the effects of which were felt in Europe and Africa as well as the rest of Asia. 

Revision of the hipparion material from Montredon, including newly excavated and other unpublished specimens brings evidence of specific heterogeneity.
A fragmentary very small MC III seems very close to H. macedonicum from the upper Vallesian of Ravin de la Pluie, Greece. In that same site was also found a large hipparion.
Most of the Montredon material is referred to H. depereti. This species associates characters usually found in Vallesian hipparions (highly plicated upper cheek teeth, deep vestíbular grooves on the lower cheek teeth, robust metapodials) with characters more frequent in Turolian forms (middle size, lack of confluence in the upper premolar fossettes, lack of ectostylids on the adult lower cheek teeth, well developped keel on the MC III, facette for the 2nd cuneiform present on all MT III). H. depereti shares some of these characters with the Spanish and Portuguese hipparions transítional between the Vallesian and the Turolian (Masia del Barbo, Azambujeira) but is not identícal to any of them. The upper Vallesian hipparion material from Diavata, Greece, probably belongs to H. depereti but not the large hipparion rests from Ravin de la Pluie.
Thus, Montredon and Ravin de la Pluie may well share the same small species, H. macedonicum, but they differ in the associated one: middle-sized H. depereti at Montredon, large-sized H. primigenium at Ravin de la Pluie. Both sites, however, give evidence of small hipparions during the Vallesian, coexisting with other larger species.
The present paper also proposes an adaptation of the Kiesewalter's indices that calculates the height at the withers using the metapodial length, and discusses indices proposed by Gromova and by Sen et al. to express the relative development of the metapodial keel. 

Résumé :
L’étude des rongeurs Theridomorpha permet de suivre le déroulement d’une radiation adaptative pendant toute sa durée (Eocène moyen-Oligocène terminal), sur un territoire restreint à l’extrémité ouest de l’Europe Occidentale. Dans ce papier, la phylogénie de ce groupe est établie à partir d’une analyse cladistique reposant sur l’examen de 315 caractères (310 dentaires). Le groupe d’intérêt comprend 110 des 132 espèces (24 genres) de Theridomyoidea et deux genres encore inclus jusqu’ici dans les Reithroparamyinae qui rejoignent les Theridomorpha. Les groupes externes comprennent des Glires basaux, Cocomys, Tanquammys et 16 Ischyromyiformes. Un cadre phylogénétique robuste est produit, qui permet de clarifier la systématique des Theridomorpha. La position des Remyoidea (nov. sup.fam.) au sein des Ischyromyiformes, extérieure aux Theridomorpha, est confortée. Les Protadelomys et Tardenomys sont à la base des Theridomyoidea, avant la séparation en deux clades correspondant aux familles Pseudosciuridae et Theridomyidae. Les sous-familles sont consolidées : Pseudosciurinae et Sciuroidinae pour les Pseudosciuridae ; Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, auxquelles s’ajoute au moins une nouvelle sous-famille (Patriotheridomyinae), pour les Theridomyidae. La topologie des chrono-espèces (sensu Simpson), traitées antérieurement comme lignées évolutives, apparaît dans la plupart des cas sous forme de clades successifs dans lesquels les espèces sont le plus souvent arrangées de manière pectinée, émergeant dans l’ordre stratigraphique. L’analyse des caractères aux principaux nœuds permet de consolider les caractères diagnostiques des taxons et les tendances évolutives, ainsi que de discuter des divers parallélismes et convergences dans l’évolution des structures et patrons dentaires (e.g., émail des incisives unisérié chez les Issiodoromyinae et les Patriotheridomyinae, ou pseudo-multisérié chez les Blainvillimys les plus hypsodontes, les Protechimys et Archaeomys ; patrons dentaires téniodontes ; allongement des dents déciduales chez les Patriotheridomyinae, Issiodoromyinae et Theridomyidae ; sélénodontie ou lophodontie). Les dynamiques évolutives traduites par les changements morphologiques sont mises en relation avec les variations environnementales. Enfin, les implications biochronologiques de l’évolution des Theridomyoidea sont discutées.
Abstract:
The adaptive radiation of the rodents Theridomorpha occurred during a limited time window (middle Eocene to late Oligocene), on an area restricted to Western Europe. In this paper, the phylogeny of this group is established via a cladistic assessment of 315 morphological characters (310 dental). The group of interest encompasses 110 upon 132 species (24 genera) of Theridomyoidea, and two genera formerly included within the Reithroparamyinae, and which are included here within the Theridomorpha. The outgroups include basal Glires, Cocomys, Tanquammys and 16 Ischyromyiformes. A robust phylogenetic frame is produced, which allows clarifying the systematics of the Theridomorpha. Within the Ischyromyiformes, the Remyoidea (nov. supfam.) are set apart from the Theridomorpha. Protadelomys and Tardenomys represent the earliest offshoots of the Theridomyoidea, before the dichotomy between Pseudosciuridae and Theridomyidae. It supports the former subfamilies Pseudosciurinae and Sciuroidinae within the Pseudosciuridae; and for the Theridomyidae: the Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, with at least one new subfamily (Patriotheridomyinae). The topologies of the chronospecies (sensu Simpson), formerly considered as evolutionary lineages, appear in most cases as successive clades, in which the species are generally pectinately arranged and emerging in the stratigraphic order. The analysis of characters supporting the main nodes allow consolidating the diagnosic characters of the taxa and their evolutionary trends, as well as discussing the various cases of parallelism and convergence in the evolution of structures and dental patterns (e.g., uniserial incisor enamel for Issiodoromyinae and Patriotheridomyinae, or pseudo-multiserial for the most hypsodont Blainvillimys, Protechimys and Archaeomys; taeniodont dental patterns; lengthening of deciduous premolars for Patriotheridomyinae, Issiodoromyinae and Theridomyidae; selenodonty or lophodonty).
Evolutionary dynamics are analysed with respect to environmental changes. Finally, biochronological implications of the evolution of Theridomyoidea are discussed.
  

The genus Plagiolophus is documented, almost solely in Western Europe, from the middle Eocene up to the mid Oligocene (MP 12 to MP 25), i.e. more than for 15 MY. Seventeen species are now recorded whose two of them are new, P. ringeadei nov. sp. and P. mamertensis nov. sp. Some anatomical variations and the deflection of certain evolutionary trends justify the distinction of three subgenera, Paloplotherium, Fraasiolophus nov. and Plagiolophus s.s. The genus displays a wide range in size and weight (between 10 and 150 kg). The detailed description of the skull of several species is here given for the first time.
Despite important evolutionary drifts during this long time span, the dentition shows a great structural homogeneity, which renders difficult the determination of fragmentary specimens or isolated teeth. It is characterized by a great heterodonty; premolars are little molarized and present a certain regression through time with paradoxically some progress in the molarization. The hypsodonty increases: the first Plagiolophus are hardly less brachyodont than Propalaeotherium, and the last ones are nearly as hypsodont as Merychippus from the early Miocene. The upper molars change from a wide crown pattern, with an open occlusal surface, lightly oblique transverse lophs and rounded internal cusps, to a narrower pattern, with a frontally constricted occlusal surface and internal lophs aligned parallel to the ectoloph. The M3/3 become always longer.

The dental enamel displays horizontal Schreger-bands with imprecise limits occupying only the middle part of the enamel layer. The dentine is remarkable by its high rate of pericanalicular dentine. The crown cementum, lacking in earlier forms, increases to the point where it fills the occlusal valleys of the
teeth.

The masticatory musculature shows a increasing prominence of the temporal, with probably an important role devoted to the pterygoid muscles in lateral movements related to a two-phase type of chewing.

The evolution of the dentition, of the masticatory musculature and of the repartition of masticatory forces indicate that the Plagiolophus have known different diets through their long evolutionary history; at first browsers they became mixed feeders and finally grazers. Their relatively long neck allowed these animals to reach different vegetal layers. The strength of the nuchal crests also suggests that they were able to have strong backwards movements of the head to pull up their food.

This evolution of diet seems related to the slow degradation of environmental conditions attested during this period in western Europe, with the generalization of more open landscapes, increasing aridity and more marked seasons.

Besides, a remodeling of the face is ontogenetically and along time observed, in relation with the evolution of the masticatory apparatus and especially with that of the mandibular lever arm. The postcanine diastemata become longer in the course of evolution; the free extremities of the nasals are always relatively long which contradicts the hypothesis according to which Paloplotherium may have had a trunk. At last the lineage Fraasiolophus can be distinguished by the presence of a deep malar fossa, probably related to a strong development of the maxillo-labialis superior muscle.

The orbit is always large and tends to increase in size, which indicates a good development of the vision and its increasing role in the life relations. A peculiar type of epitympanic sinus could have been used as a resonance chamber insuring a certain amplification of sounds before their transmission to the eardrum. The endocranial cast reveals a relatively large brain with an advanced degree of gyrencephaly. Beside the role eventually played in food research and social relations, these neurophysiological abilities, also related to an advance in cursorial fitness, could have contributed to the survival of these animals facing the predation pressure of the first fissipede carnivores and the competition with new immigrant herbivores after the "Grande Coupure".
On the basis of some shared apomorphies with the Pachynolophinae, which prevent from considering the latter as Equidae (molarization of the premolars, reduction of the premaxilla dorsal apophysis, peculiar epitympanic sinus, splitting of the jugular process), the hypothesis of an autochthonous origin of Plagiolophus issued from a form near Propalaeotherium, is once again proposed and discussed. Finally, intra-generic relationships are taken into consideration. 

No abstract available 

Biographie et liste des publications de S. Garimond. 

This paper provides a revision of the early and middle Eocene European rodents previously referred to as Ischyromyoidea, including taxa considered to be at the origin of the Theridomorpha. The use of an accurate dental terminology and a better understanding of the size and shape of their infra-orbital foramen (i.o.f.) led us to a substantial revision of this group, which allowed to better characterize them and to appreciate their variability. On these bases, phylogenetic analyses (cladistic and standard Bayesian
approaches) of early Ypresian to late Priabonian European rodent species were undertaken in order to highlight the root of the early Theridomorpha and its content. In this paper, the phylogeny was established based on 343 characters (338 dental) through 45 early Paleogene taxa using both cladistic and bayesian analyses. The ingroup included on one hand a few North American genera (Reithroparamys, Microparamys, and Acritoparamys) and European ones (Eogliravus, Ailuravus, Corbarimys, Meldimys, Euromys, Plesiarctomys, and Pseudoparamys) considered until now as being related with the North American superfamily Ischyromyoidea. On the other hand, it included genera close to the root of the Theridomorpha (Sparnacomys, Pantrogna, and Hartenbergeromys) and early Theridomyoidea (Masillamys, Protadelomys, and some Pseudosciuridae). The phylogenetic results obtained via the two
distinct reconstruction approaches are consistent in virtually all relationships. The proposed systematics here derives from these phylogenetic results. This phylogenetic context led us to change the suprafamilial, familial, subfamilial or generic attribution of several species. Characters of Theridomorpha, like the obliquely developed postprotocristid allied with the occurrence of a metalophulid I, have been found in genera previously considered as Ischyromyidae (Pseudoparamys, Euromys, Sparnacomys, Meldimys, Pantrogna, and Hartenbergeromys) as well as the large i.o.f., when preserved (Pseudoparamys, Hartenbergeromys, and Masillamys). Based on these morphological observations and new phylogenetic considerations, the content of the Theridomorpha clade is here enlarged, thereby extending back the first theridomorph radiations to the early Eocene. Aside, a new taxon (Reinomys rhomboides gen and sp. nov.) is described from Avenay. In addition, a new genus, Auroremys, is created for the species subita (Comte et al., 2012) from Chery-Chartreuve. 

This study describes four taxa of Gliridae from the Oligocene mammal locality Gröben 3: Gliravus tenuis BAI-ILO, 1975, Bransatoglis micio (MISONNE, 1957), B. planus (BAHLO, 1975) and B. heissigi n. sp. Gliravus tenuis from Gröben 3 is somewhat more advanced than the type population found in Heimersheim. This confirms previous research suggesting that Gröben 3 should be dated earlier than Heimersheim (MP 24). The first documented occurrence of B. mício around level MP 24 was found in Gröben 3. An abundance of tooth material from B. planus in Gröben 3 makes it possible, for the first time, to observe evolutionary stages within this species from MP 21 until MP 28. B. heissigi n. sp. is restricted to level MP 24. This species is located between B. mísonnei (MP 20 - 23) and Microdyromys praemurinus (MP 25 - 28). Within the lineage Bransatoglis bahloi - B. misonnei - B. heissigi, a decrease in size is noticeable.

  

A new European rodent from the middle Eocene of Spain, Zamoramys extraneus n. gen., n. sp., appears to be closely related to the middle Eocene chapattimyid rodents of Indo-Pakistan. This contradicts the generally accepted paleobiogeographic hypothesis of a Tethyian barrier between Europe and Asia isolating Europe during the middle Eocene. Because of this barrier, some authors have proposed that European and Asian rodents were not closely related, their similarity being the result of morphological convergence. Here monophyly has been tested, using the parsimony criterion, based on an analysis of dental characters (including discussing of homology and the validity of some characteristics). Our results indicate a phylogenetic relationship among the Asiatic Ctenodactyloidea, Zamoramys from Spain, and the European endemic Theridomyoidea. We also conclude from our analysis that theridomyoids and European ischyromyoids are probably not closely related phylogenetically. 

The investigations undertaken at Crivadia (Hateg Depression, Hunedoara District, Romania), the type locality of Crivadiatherium mackennai RADULESCO el al. (Radulesco, Iliesco et lliesco, 1976), led to the discovery of remains of a new Embrithopod. Close to the above mentioned species, but larger in size, this animal is here described as a new species of Crivadiatherium, C. iliescui. ln addition, the comparison made between the forms indicated above and Palaeaamasía kansui OZANSOY from the Eocene deposits of Anatolia (Ozansoy, 1966; Sen et Heintz, 1979) showed that the latter species included a heterogeneous material; this permitted us to distinguish the form in the Anatolian locality Ciçekdag-Arabin Kôyü under the name Palaeoamasia sp. The geographical distribution and diversity of the Embrithopod species under discussion (Balkan, Anatolia) support the idea of an eurasiatic origin of this group and seem to suggest the existence during the Eocene of a particular faunal province in south-eastern Europe. 

Based on isolated teeth two new scyliorhinid shark species, Scyliorhinus biformis nov. sp. and Scyliorhinus suelstorfensis nov. sp., are described from the Sülstorf Beds, early-middle Chattian, of Mecklenburg, northeastern Germany. They form part of a speciose assemblage of necto-benthic sharks and batoids which populated the warm-temperate to subtropical upper shelf sea of the south-eastern North Sea Basin. 

Based on material found in about 15 localities the relationships of the genera Microparamys and Glirarus have been studied. One new genus, two subgenera and three species [Microparamys (Sparnacomys) chandoni n. subgen. and n. sp., Microparamys (Pantrogna) russelli n. subgen., Eoglirarus wildi n. gen. and n. sp., Gliravus meridionalis n. sp.] as well as the publication
of numerous new facts concerning species previously reported, support the phyletic scheme proposed. The latter shows that the origin of the Gliravinae is to be sought among the very small and still rather poorly known Microparamys species of the early Eocene. Gliravus: robiacensis can be considered as the common ancestor to different lineages not only of Glirarus but also of modern genera (Peridyromys, Glirudinus and Microdyromyx), at the origin of which Peridyromys micio, although difficult  to interpret, occupies a similar place.

The stratigraphic conclusions permit more detail in the chronologie succession of the localities studied. The paleoecologic and biogeographic aspects lead one to the problem of the oligocene "Grande Coupure" through the study of this group.

  

The Mont-Hélène's fauna [Pyrénées-Orientales, France], includes 15 species of rodents with a new one, Occitanomys montheleni n. sp. among the 9 species of the Murids which are listed. The uncommon Cricetid, Blancomys neglectus, is well represented in the fauna. Peculiarities of the population referred to Slephanomys cf. donnezaniare discussed. The locality a fissure filling may be the oldest one of Tabianian age known in Southern France. The diversity of the Murids gives evidence of a subtropical climate and of a diversified environment which may be linked to the spreading of the coastal plain following the filling up of the Roussillon Neogene Basin. 

The rodent fauna of Beni-Mellal is characterized by the abundance of ctenodactylids and cricetids. The latter are represented by four distinct species, among which a new form. Dakkamys zaiani nov. gen., nov. sp. is described. A detailed morphological analysis shows that, contrary to that which had been established before, « Cricetodon ›› atlasi Lavocat, 1961, is not closely related to any European form known; this species is attributed, in consequence, to the new genus Mellalomys. A simple biometric analysis has shown that the genus Myocricetodon Lavocat, 1952, is represented in this locality by two distinct species. The systematic homogeneity of the Beni-Mellal cricetids is also demonstrated: they can, as a matter of fact, all be referred to the subfamily Myocricetodontinae. The definition of this subfamily is completed. The sciurids and glirids are also reviewed in the light of new systematic and biogeographic information established ln Europe. A new species of Atlantoxevus from the early Pleistocene of Morocco, A. huvelini nov. sp., is described. It is probably the descendant of A. tadlae from Beni-Mellal. Biogeographic analysis leads one to consider this fauna as the result of geographic isolation in the Maghreb since the late Oligocene or the early Miocene. In particular no direct European influence can be discerned. Stratigraphic considerations resulting from the discovery of new localities in North Africa lead to the confirmation of the ante-Vallesian age of this fauna and to its parallelism with the faunas of La Grive in Western Europe and Fort Ternan in East Africa. The peculiar geologic nature of this locality is discussed.
  

We present here a new updated counting of the perissodactyls of Robiac, the type locality of the MP 16 level of the biochronological scale of paleogene mammals and that of the Robiacian stage of Eocene Land Mammals Ages in Western Europe.
The outcrop of Robiac consists actually of two 500m apart loci, Robiac-Nord and Robiac-Sud, considered of the same age according to the current discriminating power, and is dated from -38,7 MA after the last faunal, magnetostratigraphic and climatic calibrations.
It has yielded a very abundant and rich of 21 taxa perissodactyl fauna, topped by the giant Lophiodon lautricense, last representative of the family Lophiodontidae, of which it is the last proved deposit. The Palaeotheriidae are much diversified with 5 genera and 9 species of "Pachynolophinae", 3 genera and 10 species of Palaeotheriinae. Nine taxa have been defined from Robiac: Chasmotherium depereti n. sp., Palaeotherium castrense robiacense Franzen, 1968, the genus Leptolophus Remy, 1965 with the species L. stehlini Remy, 1965 and L. magnus Remy, 1998, Anchilophus (Paranchilophus) jeanteti Remy, 2012, Metanchilophus chaubeti Remy, 2012, Lophiotherium robiacense Depéret, 1917 and Pachynolophus gaytei n. sp.
The faunal Robiac cenogram with the associated flora testify to a hot, wet and forestal environment, likely corresponding to a short warming climatic phase; the broken up fossil bones should have been carried away and then gathered in swamp areas along the banks of a meandering river.
The swarm of mammals of Robiac, the richest of contemporaneous deposits, has been followed by a drastic drop in perissodactyl diversity at the MP 17A level; a crisis which could have originated in a renewal of the global Eocene cooling. Fons 4, the type-locality of this level, is largely scarcer in perissodactyls and its cenogram testifies to a less diversified fauna, with on the whole smaller species, that likely means a cooler and drier climatic environment; a new perissodactyl diversification occurred but later.
  

     The Faunas of Rodents from seven north-african fossiliferous beds distributed from the Middle up to the Uppest Miocene are studied. One genus, seventeen species, one subspecies described are new.
       A detailed description is given of the evolution of the numerous specific lines, systematic groups and communities. The evidence of the strong endemism of the North African faunas of that time compared with those of Europe as well as of Tropical Africa is clearly shown. Special relationships with Asia clearly also appear, mostly in the Middle Miocene and the beginning of the Upper Miocene.
       Using the specific lines with a biostratigraphic purpose gives the opportunity to establish a biochronological scale for the continental formations of the Maghreb. This scale is calibrated by a few absolute datations. These datations, as well as the stratigraphical study of several fossiliferous localities have given the possibility to establish precise correlations with the corresponding scales of Europe and of Tropical Africa, as well as with the marine formations of the Neogene of occidental Mediterranean sea.
       The paleoclimatic data connected with this period in North Africa are explained and the contribution of the micromammals is discussed. 

The study of new specimens from the Stephanian shales of Montceau-les-Mines confirms and enlarges the number of groups already known in this area. Among the Palaeonisciforms, “form A" is now known more completely, although no diagnosis or name can yet be given for it. “Form B" is redescribed and its relationships with “Elonichthys robisoni" are discussed. A palaeoniscid is recorded which resembles those from Bourbon l'Archambault. The paramblypteriforms occur rather frequently, but no genera can be determined. The aeduelliforms comprise some specimens close to Aeduella blainvíllei from Muse (Autun basin), and a new genus. Comparison of the latter with two fossils from Lally allows creation of two new species and a new family. This diversification of the aeduelliforms during this middle Stephanian leads to the hypothesis that the group originated at least as early the lower Stephanian. This material prooves again the characteristic endemism of this fauna, particularly of the aeduelliforms which are known only in the Massif Central where they diversified during the Permo-Carboniferous. Biogeographical consequences are discussed. 

As a result of the enrichment of eotheriodont material by one of us (P.K.T.), these specimens (essentially Biarmosuchur and Eotitanosuchur) are reexamined and refigured. A reevaluation of their particularities supports the distinction of two families, for which new diagnoses are proposed. This leads us to discuss the affinities of these families, with respect to the sphenacodonts on one hand, and to the South African primitive theriodonts on the other (gorgonopsids and ictidorhinids). This study contains inherent paleogeographic consequences which are considered in conclusion.