Résumé :
L’étude des rongeurs Theridomorpha permet de suivre le déroulement d’une radiation adaptative pendant toute sa durée (Eocène moyen-Oligocène terminal), sur un territoire restreint à l’extrémité ouest de l’Europe Occidentale. Dans ce papier, la phylogénie de ce groupe est établie à partir d’une analyse cladistique reposant sur l’examen de 315 caractères (310 dentaires). Le groupe d’intérêt comprend 110 des 132 espèces (24 genres) de Theridomyoidea et deux genres encore inclus jusqu’ici dans les Reithroparamyinae qui rejoignent les Theridomorpha. Les groupes externes comprennent des Glires basaux, Cocomys, Tanquammys et 16 Ischyromyiformes. Un cadre phylogénétique robuste est produit, qui permet de clarifier la systématique des Theridomorpha. La position des Remyoidea (nov. sup.fam.) au sein des Ischyromyiformes, extérieure aux Theridomorpha, est confortée. Les Protadelomys et Tardenomys sont à la base des Theridomyoidea, avant la séparation en deux clades correspondant aux familles Pseudosciuridae et Theridomyidae. Les sous-familles sont consolidées : Pseudosciurinae et Sciuroidinae pour les Pseudosciuridae ; Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, auxquelles s’ajoute au moins une nouvelle sous-famille (Patriotheridomyinae), pour les Theridomyidae. La topologie des chrono-espèces (sensu Simpson), traitées antérieurement comme lignées évolutives, apparaît dans la plupart des cas sous forme de clades successifs dans lesquels les espèces sont le plus souvent arrangées de manière pectinée, émergeant dans l’ordre stratigraphique. L’analyse des caractères aux principaux nœuds permet de consolider les caractères diagnostiques des taxons et les tendances évolutives, ainsi que de discuter des divers parallélismes et convergences dans l’évolution des structures et patrons dentaires (e.g., émail des incisives unisérié chez les Issiodoromyinae et les Patriotheridomyinae, ou pseudo-multisérié chez les Blainvillimys les plus hypsodontes, les Protechimys et Archaeomys ; patrons dentaires téniodontes ; allongement des dents déciduales chez les Patriotheridomyinae, Issiodoromyinae et Theridomyidae ; sélénodontie ou lophodontie). Les dynamiques évolutives traduites par les changements morphologiques sont mises en relation avec les variations environnementales. Enfin, les implications biochronologiques de l’évolution des Theridomyoidea sont discutées.
Abstract:
The adaptive radiation of the rodents Theridomorpha occurred during a limited time window (middle Eocene to late Oligocene), on an area restricted to Western Europe. In this paper, the phylogeny of this group is established via a cladistic assessment of 315 morphological characters (310 dental). The group of interest encompasses 110 upon 132 species (24 genera) of Theridomyoidea, and two genera formerly included within the Reithroparamyinae, and which are included here within the Theridomorpha. The outgroups include basal Glires, Cocomys, Tanquammys and 16 Ischyromyiformes. A robust phylogenetic frame is produced, which allows clarifying the systematics of the Theridomorpha. Within the Ischyromyiformes, the Remyoidea (nov. supfam.) are set apart from the Theridomorpha. Protadelomys and Tardenomys represent the earliest offshoots of the Theridomyoidea, before the dichotomy between Pseudosciuridae and Theridomyidae. It supports the former subfamilies Pseudosciurinae and Sciuroidinae within the Pseudosciuridae; and for the Theridomyidae: the Issiodoromyinae, Oltinomyinae, Columbomyinae, Theridomyinae, with at least one new subfamily (Patriotheridomyinae). The topologies of the chronospecies (sensu Simpson), formerly considered as evolutionary lineages, appear in most cases as successive clades, in which the species are generally pectinately arranged and emerging in the stratigraphic order. The analysis of characters supporting the main nodes allow consolidating the diagnosic characters of the taxa and their evolutionary trends, as well as discussing the various cases of parallelism and convergence in the evolution of structures and dental patterns (e.g., uniserial incisor enamel for Issiodoromyinae and Patriotheridomyinae, or pseudo-multiserial for the most hypsodont Blainvillimys, Protechimys and Archaeomys; taeniodont dental patterns; lengthening of deciduous premolars for Patriotheridomyinae, Issiodoromyinae and Theridomyidae; selenodonty or lophodonty).
Evolutionary dynamics are analysed with respect to environmental changes. Finally, biochronological implications of the evolution of Theridomyoidea are discussed.
  

Isolated elasmobranch teeth (sharks and rays) from the middle Eocene (Bartonian) Bandah Formation in the Jaisalmer District of Rajasthan, India are described. The remains improve our knowledge of the environment represented by this lithostratigraphic unit and the ecology preserved therein. Seventeen unequivocal taxa were identified, including Nebrius sp., Striatolamia aff. S. macrota, Brachycarcharias atlasi, B. lerichei, cf. Jaekelotodus sp., Carcharhinus mancinae, Rhizoprionodon sp., Physogaleus sp., Galeocerdo clarkensis, G. eaglesomei, Odontorhytis aff. O. pappenheimi, “Rhinobatos” sp., “Dasyatis” sp., Coupatezia sp., “Aetomylaeus” sp., “Rhinoptera” sp., and Ouledia aff. O. lacuna. Of these, “Aetomylaeus” sp., B. atlasi, C. mancinae, G. clarkensis, G. eaglesomei, cf. Jaekelotodus sp., Nebrius sp., Odontorhytis aff. O. pappenheimi, Ouledia aff. O. lacuna, and “Rhinoptera” sp. are reported from the middle Eocene of India for the first time. The Bandah Formation elasmobranch palaeofauna has close affinities to the Palaeocene-Eocene Tethyan/Paratethyan faunas of Africa, Madagascar, Asia, and Europe, and some taxa indicate a western hemisphere influence from North America. The Bandah Formation palaeofauna indicates that deposition occurred in a moderately shallow marine environment. The Bartonian age is primarily based on foraminifera but is corroborated by the presence of elasmobranch taxa that also occur in contemporaneous deposits elsewhere. The marine regression started during the early Palaeogene, and our study indicates that the sea completely withdrew from the Jaisalmer Basin after the deposition of the Bandah Formation. This event may have been synchronous with the middle Eocene uplift of the Himalayan-Tibetan Plateau. 

The dentition of selachians is characterized by an often very pronounced heterodonty involving a great morphological diversity. Despite this fact, the dentitions of selachians can be grouped in a rather reduced number of dental types corresponding to trophic adaptations: grasping, tearing, cutting, crushing, grinding and grasping-grinding type. The numerous exemples of convergence and parallelism that can be observed in fossil selachians and between Recent and fossil ones is the result of this reduced number of dental types. These dental specialisations allow to try a reconstruction of the way of life of fossil forms. 

The first part of this study was devoted to a descriptive analysis of teeth morphologies among the vespertilionine bats. This leads now to a tentative synthesis, providing views on the systematics of the group. The results could be seen according to three distinct but closely related purposes : 1 - the sorting of the genera contents in order to conform the genera units to homogeneous taxa that could represent natural issues of evolutionary lineages ; 2 - the investigation of relationships between extant genera in order to infer the possibilities of common origin ; 3 - according to the preceeding items and to the observed evolutionary trends, a tentative phylogeny, modest and cautious. The contents of many genera are sorted : Leuconoe is removed from subgeneric to generic position, whereas Myotis becomes a subgenus of it ; the myotodont species are cleared away from the Pipistrellus genus ; Glischropus and Scotozous are synonymized within Pipistrellus ; Hypsugo is raised to the generic level ; some species previously ranged within Pipistrellus will form provisionally a collective group, Attalepharca nov. ; the Eptesicus genus is broken up, the excluded species being grouped within Nycterikaupius gen. nov. ; the Nycticeini tribe is defined again after exclusion of Otonycteris , Scotoecus, Scotophilus , and addition of Hesperoptenus ; the species la io and Pipistrellus tasmaniensis are removed to Eptesicus (n.s.) and Pipistrellus dormeri to Scotoecus. Groupings of genera are stated according to the main evolutionary trends of I1/. The relevance of these is often warranted by close morphologic similarities of other teeth. This leads to a recognition of the major evolutionary radiations which occurred in the group. The filiations schematized at the end of the work show the dental relationships observed between the extant genera, and could represent a phylogenic framework. Two major facts are to be underlined : 1- the early divergence of leuconoids ; 2 - the successives crossings to myotodonty from the nyctaloid flow. Fossil data from the literature are punctually and tentatively incorporated within phylogenic sketches. 

The Tafna Basin corresponds to the lowlands, which are located in front of Tessala and Traras ranges, below the Tlemcen mountains, Algeria. This basin displays a complete sedimentary cycle dominated by lagoonal-fluvial and marine deposits. The continental formations located at the base of these deposits are mainly composed of alternating sandstones and clays. An early late Miocene age has been previously attributed to them, based on direct correlations with marine deposits. Search for micromammal fossils led to the discovery of three different rodent species from a single level of the Djebel Guetaf section, located at the bottom of these deposits. The rodent assemblage indicates a late Miocene age. Combined magnetostratigraphical and biostratigraphical investigations were carried out to provide a more accurate age control of these continental deposits. Sixty-four oriented rock samples were collected for a magnetostratigraphic study along a 92 meters thick section including the fossiliferous layer. Rock magnetic investigations indicate the presence of both high and low coercivity minerals. Specimens subjected to progressive thermal demagnetization procedures show that the samples exhibit a high temperature magnetization component and display a normal polarity. Based on biostratigraphic constraints, the Guetaf section is correlated with Chron C4An, indicating an age ranging from
9.1 to 8.7 Myr.
  

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

The first lists of Insectivores (Erinaceidae, Talpidae and Soricidae) from the Pliocene beds of Southern France and North-East Spain are given in this paper. The material from twelve localities is studied. These localities are geographically situated in Languedoc (Celleneuve, Vendargues, Nîmes, Sète, Balaruc 2 and Seynes), in Roussillon (Terrats, Serrats-d'en-Vacquer, Château d'eau and Mont-Hélène) and in North-East Spain (Layna, Medas Islands and Puebla de Valverde). These faunas correspond to the Early, Middle and Late Pliocene. 1 to 8 taxa are identified in these localities and 14 specific taxa are presently listed for this period in this area. Two new specific taxa are described as Galerix depereti nov. sp. from all the Early Pliocene localities in the North-Pyrenean area and as Desmanella gardiolensis nov. sp. from Balaruc 2. For this small mammals, two faunal assemblages are recognized. The first one is dated from the Early Pliocene (F 1, 2 and 3 zones in Aguilar et Michaux) and is characterized by Galerix depereti and rare and little diversified Soricids. The second one is Late Pliocene in age (zones G 2 and G 3). The fossils of the genus Talpa are relatively abundant and the Soricids are diversified and very abundant. The Middle Pliocene (zone G 1) is a transitional period. ln these faunas, most of the insectivore genera are known from the European Late Miocene beds (8 on 10). This fact demonstrates a relative continuity between the invectivore faunas from the Late Miocene to the Early Pliocene. In conclusion, somme paleoecological considerations are suggested.
  

The dentition of Welcommia bodeuri nov. gen. nov. sp. from the Valanginian of Southem France is described and reconstructed. Species and genera of Upper Jurassic and Lower Cretaceous Hexanchiformes are reviewed and discussed.
The genus Notidanoides MAISEY, 1986 must be restricted to the single Nusplingen Upper Jurassic specimen, whose attribution to the species muensteri AGASSIZ, 1843 remains doubtful.
The genus Paranotidanus WARD &THIES, 1987 that does not rest on any type-species nor on any precise dental characterization must be rejected.
The genus Eonotidanus PFEIL, 1983, based on a very poorly preserved and heterogeneous type-material must also be rejected.
Teeth from the Lower Cretaceous, with a peculiar morphology, previously assigned to Eonotidanus or to Notidanoides, are to be ranked in the new genus Pachyhexanchus. 

Abstract
 The validity of Propachynolophus Lemoine, 1891, supposedly an intermediate between Hyracotherium Owen, 1841 and Pachynolophus Pomel, 1847, has been questioned for a long time. A detailed analysis of features on which this genus is based further supported by a formal cladistic analysis demonstrates that Propachynolophus is not a valid taxon. The type species, “Propachynolophus gaudryi Lemoine, 1891” shall be assigned to Propalaeotherium Gervais, 1849, under the new combination Propalaeotherium gaudryi (Lemoine, 1891). “Pachynolophus maldani Lemoine, 1878”, later assigned to Propachynolophus, typifies the new genus Orolophus, under the binomen Orolophus maldani (Lemoine, 1878). The other referred species, “Propachynolophus levei Hooker, 1994” and “P. remyi Checa-Soler, 1997” are poorly documented, and both species shall be provisionally referred to as “Hyracotherium” levei (Hooker, 1994) and “Hyracotherium” remyi (Checa-Soler, 1997), pending new discoveries.
 
  

Revision of ancient collections and study of new material from Provenchères-sur-Meuse (Rhaetian) lead to signíficant changes in the faunal list of this site. This bring to us important information about the effect of the rhaetian transgression on the evolution of the faunas at this period of time. However, study of
vertebrate microremains is always difficult and some points remain obscure, like the origin of the prosauropods which yield some remains at Provenchères-sur-Meuse and the survival in Europe during the Rhaetian of temnospondyl arnphibians different from the Plagiosauridae.

  

Fourteen distinct phyletical lineages which belong at least in three families: Ischyromyidae ALSTON, 1876, Gliridae THOMAS, 1896 and Theridomyidae ALSTON, 1876, have been identified after the study of more than 3600 rodent dental remains from about twenty Early and Middle Eocene european localities. A systematical and phylogenetical revision of these rodents has been achieved. Nearly all the specific and generic diagnosis are emended. Several new combinations and synonymies are proposed. Four new species and two new genera, Euromys nov. (Ailuravinae) and Hartenbergeromys nov. (Microparamyini), are named and described. Euromys nov. gen. is known by three distinctive ypresian (MP 7 to MP 10 european reference levels) chronospecies. This new lineage is thought to be the direct ancestor of Meldimys MICHAUX, 1968 and Ailuravus RUTIMEYER, 1891. A new species of the genus Plesiarctomys BRAVARD, 1850, Pl. lapicidinarum from Condé-en-Brie (MP 8-9 reference level), allows to relate the Plesiarctomys lineage to the Pseudoparamys MICHAUX, 1964 one. The taxa Sparnacomys HARTENBERGER, 1971, Pantrogna HARTENBERGER, 1971, and Corbarimys MARANDAT, 1989 are erected to genus rank; the last one is not thought to be an Ischyromyidae. A new chronospecies of Pantrogna, P. marandati nov. sp. from the locality of Prémontré (MP 10 reference level), is described. This lineage is at the origin of two others, namely Masillamys TOBIEN, 1954, including M. mattaueri (HARTENBERGER, 1975) nov. comb. (MP 10 reference level), and Hartenbergeromys nov. gen., known from MP 10 (H. hautefeuillei nov. sp.) and MP 11 (H. parvus TOBIEN, 1954) reference levels. The phylogenetical position of Hartenbergeromys nov. gen., at the origin of the european family Theridomyidae, is discussed. The systematical and phylogenetical status of two probable Paramyinae, "Paramys" woodi MICHAUX, 1964 and an unnamed genus and species, are discussed. New populations of the primitive Gliridae Eogliravus HARTENBERGER, 1971 and of the primitive Theridomyidae Protadelomys HARTENBERGER, 1968, are described and assigned to previously known species.

  

Bulk-sampling of fossil-rich tempestites from the Chattian Sülstorf Beds of
Mecklenburg, north-eastern Germany, yielded a rich selachian fauna in which batoids
predominate by the abundance of teeth but are subordinate by the number of taxa. Thirteen
taxa are identified, among which rajiform batoids are the most diverse (six species). One
genus and three species are newly described: Raja thiedei sp. nov., Oligoraja pristina gen. et
sp. nov., and Torpedo chattica sp. nov. Two species are reallocated: Atlantoraja cecilae
(Steurbaut & Herman, 1978) new comb., and Dipturus casieri (Steurbaut & Herman, 1978)
new comb. Ontogenetic heterodonty is documented for the first time in the dental pattern of
Myliobatis sp. Stratigraphical ranges of batoid taxa in the period from Rupelian to Langhian
are presented and partly discussed in context with the palaeoclimatic evolution and
palaeogeographic situation of the North Sea Basin. 

Murids and gerbillids (Rodentia, Mammalia) recovered from four Upper Siwalik localities; Moginand (around 3.5-4.5 m.y.), Kanthro (around 2.5 m.y.), Ghaggar (around 2 m.y.) and Nadah (around 1.8-2 m.y.) are described herein. A comparison of Mus linnaeusi sp. nov. reveals its close relationship towards the house mouse Mus musculus. In contradiction to earlier proposals that Golunda migrated to the Indian subcontinent from Africa sometime during Late Pliocene, it is suggested here that Golunda (in the form of Golunda tatroticus sp. nov.) evolved from Parapelomys robertsí of Late Miocene deposits of Siwaliks. Golunda tatroticus sp. nov. exhibits a progressive relationship to the extant Indian Bush Rat Golunda ellioti through Golunda kelleri and Golunda sp. Tatera pinjoricus sp. nov. is considered here to be a link between the extinct Abudhabia kabulense and the extant Tatera indica.

  

Recent collecting of fossil vertebrate remains from the lowermost member of the Aquia Formation (Paleocene), has enabled me to report here for the very fIrst time, the earliest occurrence for the teeth of Palaeocarcharodon in the fossil record of the New World.
This report represents only one species of neoselachian from this locality, the remaining fauna of which will subsequently be described. 

Many types of insectivores have been described from specimens found in the Quercy phosphorites. These remains very often were not dated because they came from old collections. Recent excavations have permitted the situation of Amphidozotherium cayluxi FILHOL in the late Eocene. Two new genera are descrlbed based on material both from the old collections (Cryplotopos nov. g.) and from that recently recovered (Darbonetus nov. g., beginning of the middle Oligocene). Their systematic positions are revised and comparisons with American faunas are made. Amphidozotherium is not a talpid, but an erinaceoid belonging to an indeterminate family. Saturninia gracilis STHELIN is classified among the Nyctitheriinae, Myxomygale antiqua FILHOL among the Urotrichini Talpinae, and the genus Geotypus POMEL among the Scaptonichini Talpinae. A new study of the talpids from Auvergne has been rendered necessary. During the late Eocene and Oligocene precise morphology relationships existed between certain insectivores of Europe and North America. 

Three erinaceomorph species are known from the early Middle Eocene of Grube Messel near Darmstadt, Germany, which are referred to the family Amphilemuridae. Pholidocercus hassiacus, Macrocranion tupaiodon, and Macrocranion tenerum showed extraordinary adaptations to their different life strategies, and several of their specializations are unknown among living insectivores. Pholídocercus was a well-defended robust animal with an opportunistic feeding strategy. Macrocraníon zupaiodon was a slender forest floor-dweller with saltatorial specializations to escape from predators; fishes were the preferred component of its omnivorous diet. Macrocranion tenerum exhibited a combination of both survival strategies, extremely elongated hind limbs for rapid and even ricochetal flight and a spiny exterior as an effective protective device; it was probably specialized for feeding on ants. Thus, closely related, omnivorous-insectivorous forest floor-dwellers could exploit the Messel ecosystem. 

Ornithocheirid pterosaurs were the largest of the toothed pterodactyloids and had a worldwide distribution, although their fossil record is fragmentary, with the exception of the north-eastern Brazilian Crato and Santana Formations (Aptian, ?Albian, Early Cretaceous). With Istiodactylidae, they were also the only toothed pterosaurs that survived into the Cenomanian (Late Cretaceous), becoming extinct at the end of this period. Here we report on an ornithocheirid metacapus from the Late Cenomanian laminated limestone of north-eastern Mexico discovered about 120 km north-west of Ciudad Acuña, northern Coahuila at the south banks of Rio Bravo. The specimen comprises a fragmentary distal syncarpal, a crushed but complete metacarpal IV, two fragmentary preaxial metacarpals and a possible fragmentary terminal left wing finger phalanx. It represents the geologically youngest known ornithocheirid worldwide. We suggest that ornithocheirid pterosaurs may have become extinct because of massive sea level fluctuations during the mid to late Cretaceous that may have obliterated their breeding sites on coastal plains and low lying islands. 

A new carcharhinid shark species, Physogaleus hemmooriensis sp. nov., is described from the Lower Hemmoorian (Behrendorfian, late Burdigalian, early Miocene) of Werder, Lower Saxony, Germany. P. hemmooriensis also occurs in the Edegem and Antwerpen Sands Members of the Berchem Formation, Belgium, and in the Miste Bed, Aalten Member of the Breda Formation, The Netherlands, which have an early to middle Miocene age. In the Western Atlantic region, the taxon is present in the early Miocene Calvert Formation of Delaware, U.S.A, which is largely contemporaneous with the Hemmoorian. 

The Mont-Hélène's fauna [Pyrénées-Orientales, France], includes 15 species of rodents with a new one, Occitanomys montheleni n. sp. among the 9 species of the Murids which are listed. The uncommon Cricetid, Blancomys neglectus, is well represented in the fauna. Peculiarities of the population referred to Slephanomys cf. donnezaniare discussed. The locality a fissure filling may be the oldest one of Tabianian age known in Southern France. The diversity of the Murids gives evidence of a subtropical climate and of a diversified environment which may be linked to the spreading of the coastal plain following the filling up of the Roussillon Neogene Basin. 

The classifications of the recent vespertilionine bats were made wihtout taking in account the teeth morphology; this resulted in a reduction of the possibilities of comparison with the available fossils. The generalized use of dental formulae was abusive: this contributed to the admission of artificial genera. These conditions have long delayed the consideration of characters able to frame the phylogeny of the sub-family. In the first part of the study, the teeth morphologies are described and analysed. morphological reference types are established for each upper and lower tooth: they should make an easier elaboration of criteria for the differentiation at generic level. The position of the species in view of these criteria allows one to group them into homogeneous genera, and to appreciate the degree of relationship that the latter have between them. The second part of the study (next publicationà will develop inferences dealing with systematics and phylogeny