A strong lophodonty, an extreme heterodonty, some hypsodonty and regular overlayings of coronal cement are prominent features of the genus Leptolophus (Palaeotheriinae = Palaeotheriidae s.s.). The histological pattern of the teeth unusually joins type II enamel prisms, characteristic of advanced ungulates, together with archaic features, such as an almost complete lack of Hunter-Schreger zonation and a weak expanse of peritubular dentine. The skull is narrow and slender, with an elongated ante-orbital facial region, a moderately notched nasal aperture, a rather elongated post-canine diastem, parallel zygornatic arches and a fairly dorsally located squamoso-mandibular joint.The functional analysis brings to light "ectolophodont" masticatory cycles with two phases, in which maximum power was applied, contrary to equíds, on hindmost teeth; likewise, skull accomodations to increasing height of the teeth are quite different. This study leads to the assumption that Leptolophus may have been light mammals, living in rather open surroundings, browsing on herbaceous plants or leaves cropped close to the ground. Moreover, it appears that it could have been some inadequacy of dental structures to the dietary, which leaded to quick wear of the teeth and to many enamel notches, but had been somewhat balanced by the early increase of hypsodonty, not induced in such a case by a biotop deterioration (as it will happen at the end of the Eocene). This ínadaptation might account for the short duration of the genus Leptolophus, whose the 3 species, L. stehlini, L. nouletí and L. magnus n. sp. are indeed confined in the level MP 16. Its geographical spreading (as far as known, South of western Europe) and the morphological pattern of its dentition suggest that this genus would have been related to early upper Eocene endemic spanish forms.





  

The revision of the old collections of fossil birds from the “Phosphorites du Quercy” and the study of new material give the following results (Gruiformes, Cariamae) :  The humeri and most of the carpometacarpi described under the name Filholornis belong in Elaphrocnemus. The ulnae ascribed to Fïlholornis belong in Idiornis. Most of the post-cranial elements of the genera Elaphrocnemus and Idiornis are described and show great similarities with recent Cariamidae and Opisthocomidae, and fossil Bathornithinae.  A new genus and a new species, Oblitavis insolitus, are created in the sub-family Idiornithinae; two new species are described in the genera Elaphrocnemus (E. brodkorbz) and Idiornis (I. itardiensis), and the species Elaphrocnemus gracilis is transferred to the genus Idiornis.  The genus Propelargus Lydekker is transferred from the family Ciconiidae to Cariamidae.  A new generic name, Occitaniavis, is created for the species Geranopsis elatus, which belong in Cariamidae, while the type-species of the genus, Geranopsis hastingsiae, is a member of the Gruidae.  The affinities between the Quercy avifauna and the Neotropical one is emphasized by the occurrence of Phorusrhacidae, previously known only from the Cenozoic of South America and the Late Pliocene or Early Pleistocene of North America. Thanks to the material collected during the new excavations, the stratigraphical position of most of the species is stated precisely, and evolutionary lineages are outlined. This study shows that the suborder Cariamae, presently restricted to two South American genera, was already extremely diversified during the Eocene, and widespread in Europe and North America. 

La poche de Ste-Néboule, commune de Béduer (Lot), 15 km environ à l'WSW de Figeac, fait partie du groupe le plus septentrional des gouffres creusés par les ruissellements du Paléogène dans les calcaires jurassiques de la bordure sud-ouest du Massif Central et qui furent comblés à la même époque par des argiles sidérolithiques accompagnées de phosphate de chaux concrétionné ainsi que des restes de la célèbre faune dite «des phosphorites du Quercy» . 

A new Propalaeotherium species, clearly distinct from the genus Eurohippus, is described. It is characterized by having a similar size as P. voigti from the German Geiseltal localities (MP 11 to MP 13 reference-level), but differs in several features suggesting a slighty more derived morphology. It presents indeed less brachyodont crowns with less prominent and less elevated cingula, slightly larger relative surface of premolars, and a more marked metaconid splitting on cheek teeth. This new species is unknown from other European localities except the nearby Saint-Martin de Londres locality which has been considered older than the MP 13 level. 

Morphological and biometrical description of skulls, teeth, and limb bones of extant and fossil Old World herniones (including E. hydruntinus) and of New World 'stilt-Iegged' and other slender species from Blancan to Holocene. An Appendix presents ways in which the approximate size of some missing bones or dimensions may be deduced from available ones.

The discussed and/or illustrated fossils were found in Bolivia (Tarija), Canada (Yukon), China (Choukoutien, Gulongshan, Jiling, Loufangzi), Ecuador (Oil Fields), Ethiopia (Melka Kunturé), France (Lunel-Viel), Germany (Süssenborn), Greece (Agios Georgios, Petralona), Hungary (Dorog), Italy (Romanelli), Mexico (Cedazo, San Josecito), Mongolia (Sjara-osso-gol), Spain (Venta Micena), ex-Soviet Union (Akhalkalaki, Binagady, Chokurcha, Chukochya, Kabazi, Kolyma, Krestovka, Kurtak, Staroselie, Tologoj), USA (Alaska, Arkalon, Cedar Meadow, Channing, Conkling, Dry Mountains, Hay Springs, Leisey Shell Pit A, Lissie Formation, Natural Trap, Pool Branch, Powers Ranch, Rock Creek, San Diego, Santo Domingo, Seymour Formation, Shelter, Slaton, Trinity River). Numerous raw or statistically elaborated data are given in Tables.

There is no evidence for the existence of Old World hemiones in the New World nor of 'stilt-Iegged' equids in the Old World. The first 'stilt-Iegged' equid was found at Santo Domingo, New Mexico, and is believed to be Late Blancan. It was probably at the origin of E. calobatus (Arkalon, Rock Creek) and of the smaller E. semiplicatus (Channing, Rock Creek). Slender, but not 'stilt-Iegged', equids found at Natural Trap, Wyoming, ca. 12 ky ago belong to Amerhippus. AlI these species share with Oid World Sussemiones (and some hemiones) peculiar patterns on the lower cheek teeth.

The slender Equus sp. B of Leisey Pit A, Florida, ca. 1.2 Ma, as weIl as Amerhippus francisci and E. tau (probably a senior synonym of E. quinni) share conventional lower cheek teeth patterns. The skulls of A. francisci and E. tau, however, are quite different.

Paleontological data suggest a common origin of Amerhippus, Sussemiones, and 'stilt-Iegged' equids during the late Blancan. Old World hemiones seem to have differentiated later. 

Many types of insectivores have been described from specimens found in the Quercy phosphorites. These remains very often were not dated because they came from old collections. Recent excavations have permitted the situation of Amphidozotherium cayluxi FILHOL in the late Eocene. Two new genera are descrlbed based on material both from the old collections (Cryplotopos nov. g.) and from that recently recovered (Darbonetus nov. g., beginning of the middle Oligocene). Their systematic positions are revised and comparisons with American faunas are made. Amphidozotherium is not a talpid, but an erinaceoid belonging to an indeterminate family. Saturninia gracilis STHELIN is classified among the Nyctitheriinae, Myxomygale antiqua FILHOL among the Urotrichini Talpinae, and the genus Geotypus POMEL among the Scaptonichini Talpinae. A new study of the talpids from Auvergne has been rendered necessary. During the late Eocene and Oligocene precise morphology relationships existed between certain insectivores of Europe and North America. 

Abstract not available 

A new genus, Eppsillycteris, is erected for Adapisorex? allglicus COOPER, 1932, from the earliest Eocene Blackheath Beds of Abbey Wood, London, England. Various derived character states indicate that it belongs to the order Chiroptera (bats) rather than to the extinct "insectivore" family Adapisoricidae. Other derived character states are shared with fossil and modern members of the family Emballonuridae. Placement of the new genus in this family extends the record of the Emballonuridae back in time by about 10 million years. It is the earliest record of a modern bat family and one of the earliest bats. This implies that the differentiation of at least some modern bat families took place in the Palaeocene, where no authenticated records of bats yet exist. The primitive characters of the earliest bats make the family Nyctitheriidae an unlikely stem group for the order Chiroptera. A tentative plausible alternative exists in some unnamed upper molars from the Palaeocene of Walbeck, Germany. Wyollycteris chalix, described as a bat from the Late Palaeocene of Wyoming, U,S.A., fits better in the family Nyctitheriidae. 

Two new finds of pantodont materials from southern China, assigned to Archaeolambda, are described in this paper. One, a new species from the Nanxiong Basin, Guangdong Province, is similar to Alcidedorbignya inopinata from the early Paleocene of Tiupampa, Bolivia in size. It provides reliable evidence of the occurrence of Archaeolambda in the early-middle Paleocene of southern China. The second find includes specimens of Archaeolambda sp. cf. A. planicanina from the ?late Paleocene of Hengyang Basin, Hunan Province, which are the first record of a fossil mammal from the area near Hengyang city. The only vertebrate fossils previously found here were two genera of crocodiles discovered in 1938. This find sheds new light on the local biostratigraphy. 

We present here a new updated counting of the perissodactyls of Robiac, the type locality of the MP 16 level of the biochronological scale of paleogene mammals and that of the Robiacian stage of Eocene Land Mammals Ages in Western Europe.
The outcrop of Robiac consists actually of two 500m apart loci, Robiac-Nord and Robiac-Sud, considered of the same age according to the current discriminating power, and is dated from -38,7 MA after the last faunal, magnetostratigraphic and climatic calibrations.
It has yielded a very abundant and rich of 21 taxa perissodactyl fauna, topped by the giant Lophiodon lautricense, last representative of the family Lophiodontidae, of which it is the last proved deposit. The Palaeotheriidae are much diversified with 5 genera and 9 species of "Pachynolophinae", 3 genera and 10 species of Palaeotheriinae. Nine taxa have been defined from Robiac: Chasmotherium depereti n. sp., Palaeotherium castrense robiacense Franzen, 1968, the genus Leptolophus Remy, 1965 with the species L. stehlini Remy, 1965 and L. magnus Remy, 1998, Anchilophus (Paranchilophus) jeanteti Remy, 2012, Metanchilophus chaubeti Remy, 2012, Lophiotherium robiacense Depéret, 1917 and Pachynolophus gaytei n. sp.
The faunal Robiac cenogram with the associated flora testify to a hot, wet and forestal environment, likely corresponding to a short warming climatic phase; the broken up fossil bones should have been carried away and then gathered in swamp areas along the banks of a meandering river.
The swarm of mammals of Robiac, the richest of contemporaneous deposits, has been followed by a drastic drop in perissodactyl diversity at the MP 17A level; a crisis which could have originated in a renewal of the global Eocene cooling. Fons 4, the type-locality of this level, is largely scarcer in perissodactyls and its cenogram testifies to a less diversified fauna, with on the whole smaller species, that likely means a cooler and drier climatic environment; a new perissodactyl diversification occurred but later.
  

Confident attribution of bipedal tridactyl dinosaur tracks to theropods or ornithopods can be challenging. Here we describe trackways produced by tetanuran dinosaurs, previously attributed to hadrosaurs, from Coahuila State, northeastern Mexico. Multiple trackways headed in the same direction suggest gregarious behaviour in these late Campanian theropods.  

The rodent fauna of Beni-Mellal is characterized by the abundance of ctenodactylids and cricetids. The latter are represented by four distinct species, among which a new form. Dakkamys zaiani nov. gen., nov. sp. is described. A detailed morphological analysis shows that, contrary to that which had been established before, « Cricetodon ›› atlasi Lavocat, 1961, is not closely related to any European form known; this species is attributed, in consequence, to the new genus Mellalomys. A simple biometric analysis has shown that the genus Myocricetodon Lavocat, 1952, is represented in this locality by two distinct species. The systematic homogeneity of the Beni-Mellal cricetids is also demonstrated: they can, as a matter of fact, all be referred to the subfamily Myocricetodontinae. The definition of this subfamily is completed. The sciurids and glirids are also reviewed in the light of new systematic and biogeographic information established ln Europe. A new species of Atlantoxevus from the early Pleistocene of Morocco, A. huvelini nov. sp., is described. It is probably the descendant of A. tadlae from Beni-Mellal. Biogeographic analysis leads one to consider this fauna as the result of geographic isolation in the Maghreb since the late Oligocene or the early Miocene. In particular no direct European influence can be discerned. Stratigraphic considerations resulting from the discovery of new localities in North Africa lead to the confirmation of the ante-Vallesian age of this fauna and to its parallelism with the faunas of La Grive in Western Europe and Fort Ternan in East Africa. The peculiar geologic nature of this locality is discussed.
  

The recent excavation of the Upper Miocene locality of Montredon (Hérault) provides a rich material of Mammals. The present study deals with rodents. The new quarry is 10 meters above the classical Depéret's Quarry. No differences have been noticed between the rodent populations coming grom these two sites. The description of two new species : Hispanomys mediterraneus and Muscardinus heintzi is given and the dormouse Eliomys is recorded there for the first time. The new zonation of the Miocene given by Aguilar (1982) and the previously established correlations between marine and continental chronological scales give Tortonien age (estimated between 9 and 10 M.Y. old) to the site of Montredon, which belongs to the Mammal zone D 2. 

A detailed comparative study of a complete skeleton of Bachitherium and a cladistic analysis of the sub-order Neoselenodontia lead us to propose a cladogram and a new classification of this group. The Tylopoda are the sister-group of the Ruminantia, which are chiefly defined by the fusion of the cuboid and navicular. Within this infra-order, Amphimeryx is the sister genus of a tetraselenodont group, in which the Hypertragulidae are well-separated group from a monophyletic group defined by the loss of trapezium, fusion of capitatum and trapezoid, and the isolation of the hypoconid on lower molars. The most primitive genera of this group, Lophiomeryx and Iberomeryx still have an open trigonid on the lower molars, but this is lingually closed in Archaeomeryx, sister-genus of the higher Ruminantia which have fused metatarsals and more evolved milk teeth. We divide them into two pan/orders : Tragulina (including the recent and miocene Tragulidae, and the North-American Leptomerycidae), and Pecora, with reduced lateral metacarpals and a new crest (telocristid) on the lower premolars. Within the Pecora, the upper molars of Gelocus are more primitive than those of Bachitherium (a genus with many autapomorphies in the dentition) itself more primitive than the group Prodremotherium + Eupecora, with fused metacarpals. We consider the Eupecora (including several genera without frontal appendages) to be monophyletic. 

Paleosols and the taphonomy of vertebrate fossils in the Eocene Kuldana Formation of northern Pakistan provide important information on the preservation and time-averaging of fossil assemblages. Morphologic, mineralogic, and chemical data as well as comparisons with Quaternary soils suggest that Kuldana paleosols formed under generally dry and oxidizing conditions over time intervals of less than 100 000 years and perhaps as short as 1000 years. The distribution of carbonate in Kuldana paleosols further indicates that the upper half of the profiles were acidic whereas the lower halves were alkaline. Vertebrate fossils are rare in Kuldana paleosols and occur primarily in well-cemented sandstones and conglomerates with abundant micritic and iron-stained nodules that were reworked from floodplain soils. The scarcity of vertebrate remains in Kuldana paleosols probably reflects a combination of acidic, dry, and oxidizing conditions in the upper half of the profiles and rapid floodplain sedimentation. Comparisons between the taphonomic characteristics of Kuldana channel fossil assemblages and bone accumulations in modem rivers provide a basis for estimating the length of time represented by Kuldana fossils from several important localities. Vertebrate fossil assemblages from Barbora Banda are characterized by a low-diversity paleofauna, partially articulated skeletons, and bones that are sorted by size and shape. Comparison with bone accumulations in modern rivers suggests that the fossils from Barbora Banda accumulated in 1 to 10 years. Vertebrate fossils from the Lower Kuldana in the Kala Chitta Hills region, typified by locality H-GSP 62, are characterized by a high-diversity paleofauna and generally random and unsorted fossil bone distributions, which suggest that the fossils from these localities represent longer time intervals than the Barbora Banda fossils. Based on the time estimates for Kuldana paleosol development, fossil assemblages in Kuldana channel deposits in the Kala Chitta Hills region probably represent time intervals of about 1000 years. 

Three erinaceomorph species are known from the early Middle Eocene of Grube Messel near Darmstadt, Germany, which are referred to the family Amphilemuridae. Pholidocercus hassiacus, Macrocranion tupaiodon, and Macrocranion tenerum showed extraordinary adaptations to their different life strategies, and several of their specializations are unknown among living insectivores. Pholídocercus was a well-defended robust animal with an opportunistic feeding strategy. Macrocraníon zupaiodon was a slender forest floor-dweller with saltatorial specializations to escape from predators; fishes were the preferred component of its omnivorous diet. Macrocranion tenerum exhibited a combination of both survival strategies, extremely elongated hind limbs for rapid and even ricochetal flight and a spiny exterior as an effective protective device; it was probably specialized for feeding on ants. Thus, closely related, omnivorous-insectivorous forest floor-dwellers could exploit the Messel ecosystem. 

Previous chronological arrangements of South American Quaternary land mammal faunas are appraised on the basis of current geological and paleontological data. Three South American late Pliocene-Pleistocene land mammal ages are conventionally recognized, from oldest to youngest, the Uquian, Ensenadan, and Lujanian ; all are defined on Argentine faunas.

     The Uquian is based fundamentally and historically on the fauna from the Uquía Formation in Jujuy Province, northwestern Argentina. Important known formations in Argentina yielding Uquian Age faunas include the sub-surface Puelche Formation (or Puelchense) near the city of Buenos Aires, and the Barranca de Los Lobos and Vorohué Formations between Mar del Plata and Miramar, Buenos Aires Province. A tentative subdivision is propos-ed for the Uquian into three subages based on knowledge of the Mar del Plata-Miramar sequence, from oldest to youngest, the Barrancalobian, Vorohuean, and Sanandresian. In Argentina the Uquian is presently marked by the first known record of Scelidodon, Hydrochoeropsis, Ctenomys, Canidae, Ursidae, Gomphotheriidae, Equidae, Tapiridae, Camelidae, Cervidae, and the last known record of Thylatheridium, Thylophorops, Dankomys, Eumysops, Pithanotomys, Eucoelophorus, Hegetotheriidae, Sparassocynidae, and Microtragulidae.

The Ensenadan Age is based on the fauna from the Ensenada Formation near the city of Ensenada, Buenos Aires Province. In Argentina the Ensenadan is marked by the first known record of Lomaphorus, Neothoracophorus, Plaxhaplous, Cavia, Lyncodon, Lutra, Galera, Smilodon, Dicotyles, Lama, Vicugna, the last known record of Orthomyctera, and the only known record of Brachynasua.

     Typícal beds of late Lujanian Age in Argentina consist of fluvial deposits occupying stream channels, and shallow basins, often incised into beds of early Lujanian (i.e. Bonaerian of early workers) and Ensenadan Age. The Lujanian Age is based on a fauna from beds along the Rio Luján, about 65 km west of the city of Buenos Aires, Buenos Aires Province. The Lujanian in Argentina is marked by the first record of Equus, Chlamyphorus, and Holochilus, and the last record of Megatherioidea, Glyptodontoidea, Arctodus (=Arctotherium), Smilodon, Litopterna, Notoungulata, Proboscidea, Equidae, Morenelaphus, and Palaeolama.

   These land mammal ages are often difficult to recognize in other South American countries. The compositions of South American Pleistocene faunas vary with the environment. Some taxa were widely distributed in fossil deposits throughout the continent, but their occurrences need not reflect synchroneity. This is a result of changing climates and habitats in time. Consequently, proposed intracontinental correlations need confirmation based on magnetostratigraphy and a radioisotope time scale. Paleontologic characterizations of these land mammal ages (i.e. first and last record, and guide fossils) are useful for much of Argentina, but extensions to most of the other parts of South America are at best tenuous.

The majority of known non-Argentine Pleistocene faunas are believed to be Lujanian in age. Possible non Argentine early Pleistocene (Uquian) faunas include Ayo Ayo and Anzaldo in Bolivia, and Cocha Verde in southern Columbia. A possible middle Pleistocene (Ensenadan or early Lujanian) fauna is the Chichense of Ecuador. Paleomagnetic and radioisotopic date (MacFadden et al., 1983) clearly indicate that the greater part of the Tarija fauna (Bolivia) is Ensenadan in age.

  The end of the Pleistocene and beginning of the Holocene in South America is marked by extinction of nearly all large mammalian herbivores and their specialized large predators. Radiocarbon age determinations suggest that large scale extinctions of megafauna occurred between 15,000 and 8,000 yrs. B.P. (years before present). 

The skull remains referred to the genus Palaeotherium are recorded and described. Biometrical tests are made to elucidate intrageneric allometric relationships and to allow comparisons with various other perissodactyls. Apart from the well known shortness of post canine diastems and deepness of the narial opening, the genus is characterized by a great lengthening of the splanchnocranium, owing to a spreading of the post-orbital facial region, by a reduced area of the eye-socket and by the prevalence of the temporal muscle with regard to the masseter; this original shape of the masticatory apparatus needs to be related to the morphology of the jugal teeth and particularly to their  asymmetrical semi-hypsodonty.
    These animals, whose running ability was evidently poor, appear to have been adapted to rather closed environments, feeding on relatively soft vegetable matter; olfactory sense was likely to play a leading part in interindividual and environmental relationships. Such evolutionary trends might explain the disappearance of most of them, as clirnatic conditions deteriorated at the end of the Eocene, before the "Grande Coupure" which affected mammalian faunas at that time.
      Although the present paper is not directly concemed with phylogenetics, it invalidates the supposed ancestor-descendant relationship between P. castrense and P. magnum, and it suggests a possible emergence of the P. medium lineage from a P. siderolithicum stock. Moreover, the structure of the post-orbital facial area allows the  erection of a new sub-genus, Franzenitherium, for the  species lautricense and duvali.

  

The Laprade fauna is chronologically situated between those from Egerkingen and Lissieu and consequently, is close to the MP 14 reference-level of the European mammalian biochronological scale (Symposium of Mainz, 1987).
This new fauna is presently the oldest known in the Quercy phosphorites, formerly the Le Bretou fauna (MP 16) was considered as the oldest one. The Laprade fauna includes 21 species which belong in 7 mammalian orders (Marsupialia: Amphiperatherium bastbergense, Amphiperatherium goethei; Apatotheria: Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp.; Lipotyphla: Saturninia cf. mamertensis, Saturninia cf. intermedia; Chiroptera: Vespertiliavus lapradei nov. sp.; Rodentia: Protadelomys cf. lugdunensis, Elfomys nov. sp.; Primates: Nannopithex cf. filholi, cf. Pseudoloris or Pivetonia; Perissodactyla: ?Palaeotherium ?castrense, small-sized Palaeotherium sp., Plagiolophus sp., Anchilophus sp.; Artiodactyla: Dichobune cf. robertiana, Mouillacitherium cartieri, Tapirulus cf. depereti, Mixtotherium priscum, Pseudamphimeryx schlosseri, and Artiodactyla indet). Sixteen of these species are mentioned for the first time from the Quercy faunas.
The recognition of a new apatemyid, Heterohyus (Gervaisyus) pygmaeus nov. subgen., nov. sp., attests to an early origin of a lineage known in the Late Eocene. The emballonurid bat Vespertiliavus lapradei nov. sp. is presently the earliest record of this genus and family. This Auversian fauna leads to discuss the age of taxa showing archaic features. These taxa were defined on specimens collected in the Quercy during the last century and have never been found in the Quercy localities recently investigated. This fauna bears also evidence of a karstic filling episode older than those previously dated by fossils in the Quercy Jurassic. 

Since the 19th century, thousands of footprints were observed in the geological series of the French Mesozoic. All are located in the Triassic and Jurassic. After a promising beginning, in France, it is only a few papers which will be published in the first half 20th century, unlike the USA and of others countries of Western Europe. One ought to wait about 1950 for a revival and now they are nearly 200 papers which were devoted to the ichnofossils. The literature abundance and the renewed interest of the naturalists for the palichnologic studies decided to us to write a synthesis work. This one begins with a stratigraphic inventory in which, localisation, age and paleontological contents of about 180 fossiliferous sites are specified. After having pointed out the followed methods, the footprints paleontological interpretation is then approached in detail and the results obtained are replaced in stratigraphy to deduce the fauna evolution during the Mesozoic. So, it appears that Ichnologic data, more varied and rich in the Triassic and Liassic than those relating to the bones, very rare for the considered periods, are very informative. The middle Triassic (Anisian-Ladinian), thus reveals Cotylosauria, Lepidosauria, Crurotarsi with Rauisuchia, Ornithosuchidae, Crocodylia and Dinosauromorpha more the "Prodinosauria": Dinosamiforme whose skeletons are known in Argentina but only in Ladinian. The rather fast domination of Dinosaurs during Norian is also as well shown. The almost exclusive presence of their footprints, up to fifty cm long, in the Lower Hettangian indicates their supremacy in the environments. Footprints characterise not very deep life places located between inter-supratidal limits and often out of water. Sedimentologic and Palaeontologic studies showed that they were great coastal spaces during Middle Triassic, flood-plain with sebkhas while Upper Triassic, and a large !!coastal marsh!! in Grands-Causses during Liassic in which, mainly, fine stromatolithic layers were deposited. During the same periad, bay beaches spread in Vendée. During the Middle Jurassic, they are also brackish to lacustrine environments and recifallagoons in- the Upper Jurassic. Numerous measurements of the footprints and trackways directions showed that the animaIs moved there in weil defined directions, for long periods. They seem due to the palaeotopography of the life environments relatively stable. Also, the discovery of vegetal radicular networks and small footprints far away from the continental borderlands has suggested that the animals continuously lived in these palaeoenvironnements, belonging to large ecosystems, where the sedimentation rate was weak. This explains that thebadies could not fossilize there but only their footprints through the cyanobacterian action in main cases. From the vertical distribution of different ichnospecies, defined with adapted statistical methods, explained in this work, a palichnostratigraphy was established for the Middle Triassic. Although the footprints are also abundant in Hettango-Sinemurian of "Grands-Causses" and the Vendée, it was not possible, up to now, to establish any zonation in this series; Probably because the palichnofauna is too little diversified there, currently reduced to a majority of Theropods II-IV tridactyl traces.