Two new species, Omanodon minor n. g., n. sp. and Shizarodon dhofarensis n. g., n. sp., known from fifteen isolated teeth, are described here as the first adapiform primates (?Anchomomyini) recognizable to date in the Taqah material (early Oligocene of Sultanate of Oman).

Omanodon minor n. g., n. sp. displays special morphological similarity to the adapid tribe Anchomomyini from the Eocene of Europe, and especially to the Anchomomys lineage. Resemblances with the extant lemurifonn Microcebus are also noticeable and could be regarded as supporting Schwartz & Tattersall (1983) hypothesis of special relationships between the anchomomyine adapids and the cheirogaleid lemuriformes. However, these morphological affmities can be interpreted, altematively, as the results of parallelisms: important differences in upper molars indicate that the resemblances of cheirogaleids and Omanodon minor n. g., n. sp. are indeed probably due to parallelisms. Phyletic relationship of O. minor n. g., n. sp. to Anchomomyini is finally the most likely hypothesis.

Shizarodon dhofarensis n. g., n. sp., although much more poorly known, is closely related to Omanodon minor n. g., n. sp., at least at a familial level. The general morphology of this species suggests
also a close link with adapid Anchomomyini, although precise relationships within this tribe remain obscure. Interesting resemblances of Shizarodon dhofarensis n. g., n. sp. to Djebelemur martinezi lower molars (early Eocene of Tunisia) are also noticeable. These resemblances are even stronger than those betwen Omanodon minor and Djebelemur martinezi. However the very bunodont upper molars referred to D. martinezi are unusual for adapids, and there are moreover some notable differences in their lower molars. Thus resemblances in Djebelemur and Shizarodon are probably due to paralellisms.

Because of the fragmentary nature of the material and of possible parallelisms, the systematic position of Omanodon and Shizarodon within adapiformes cannot however yet be established definitively. 

Afin d'étudier la magnétostratigraphie de la colline de Montredon, j'ai prélevé en Avril 1982, 93 carottes orientées réparties sur 29 sites. J.-P. Aguilar et I. Loridant m'ont aidé durant cette opération. Pour la stratigraphie de la coupe échantillonnée et pour le contenu faunique des deux gisements de mammifères de cette colline, voir J.-P. Aguilar & J.-Y. Crochet (1982).
[...] 

A general view is given of the vertebrate faunas, Cretaceous to Quaternary of age, found in Columbia and of their principal characteristics. This view leads to the discussion of the isolation of the South American continent and of the role played by the Bolivar syncline with respect to North American immigrants during the Oligocene. The absence of marine deposits of Oligocene age in the north and northwest of Columbia suggests the possibility of a communication with Central America. This communication would have permitted the passage of hystricomorph rodents, of platyrrhine monkeys, and of colubrids. The non-occupation, until then, of the ecologie niches of these groups would have favored their installation beside the indigenous fauna. In this hypothesis it would no longer be necessary to admit that these vertebrates arrived as «island hoppers ››. The eco-biologic conditions would explain the absence of large-sized forms of North American origin. 

The comparative morphology and biometry of the lower cheek teeth of modern Equus are studied on approximately 300 mandibles belonging to the 10 usually recognised species : Equus grevyi, E. burchelli, E. quagga, E. zebra, E. africanus, E. asinus, E. hemionus, E. klang, E. przewalskii, E. caballus. The studied parameters comprise : occlusal length and width, postflexid length and index ; shape of the double knot (metaconid + metastylid + lingual groove) ; depth of the vestibular groove on the molars ; frequency of the pli caballinid, protostylid and other enamel plications or islets ; frequency of the dP/l.
The same methods of study are applied to a number of North American, Eurasian and African species. For the sake of comparison, some Hemphillian equids were observed (Dinohippus interpolatus, Dinohippus leidyanus, Astrohippus ansae, Phiohippus mexicanus) but most of the discussed material belongs to Pliocene or Pleistocene species of Equus : the « stenonine ›› E. stenonis, E. simplicidens, E. sanmeniensis and E. teilhardi; the « caballine ›› E. scotti, E. lambei, E. Iaurentius, E. mosbachensis, E. germanicus, E. gallicus, E. taubachensis and the Liakhov horse. The relationships of other species, in particular the North American E. calobatus, E. occidentalis, E. cf mexicanus are not clear for the moment. ln Africa, the Plio-Pleistocene species from Koobi Fora (Kenya) show some stenonine and perhaps asinine affinities. The relationships of E. numidicus and E. tabeti are uncertain but these species are probably related to the East African ones. E. mauritanicus is most certainly related to the Quagga group.
The biometrical data are gathered in 32 tables ; 4 photographie plates and 19 figures illustrate the next. The whole is a complement of the previously published studies of the skulls, upper cheek teeth, incisors and metapodials of modern and fossil Equus.

  

Abstract not available 

The Zanclian marls from Papiol provided otoliths belonging to 53 teleost taxa; nine of those are new for the Mediterranean Pliocene. The association reflects a bathymetry between 150 and 350 m, but it is likely that such depths only existed at the initial stage of flooding of the Llobregat bay. A compilation of the available data for the whole Mediterranean realm at Zanclian time provided a list of 163 taxa, of which 105 could be identified at species level. This last group provides the most useful data for evaluating the composition and affinities of the Mediterranean Zanclian fauna, which appears to be significantly different from the Recent Mediterranean fauna. The Zanclian fauna counts 68 % of Recent species: 12,5 % were already represented in the Mediterranean Miocene, 37 % appeared there at the Zanclian, and 18 % are extra-Mediterranean today. This last group is essentially composed by oceanic fishes, living in the warm parts of the Atlantic and Indo-Pacific Oceans. It is evident that a non neglectable number of Recent species only penetrated very recently in the Mediterranean, which stresses the differences between the more oceanic Zanclian Mediterranean fauna and the present one.


  

Rodents from the new localities of Châteauredon (Alpes-de-Haute-Provence) and La Denise (Bouches-du-Rhône) show more similarities with the rodents known from the Czecho-Slovak locality of Franzenbad than with the species found in contemporaneous Lower Miocene faunas located westward of the present Rhône Valley. This is another data wich support the existance of several biogeographical provinces in Southern France during Lower Miocene. The correlations which have been settled down allow new datations of several formations in Provence and give new information on the Aquitanian and Burdigalian paleogeographies of this region. More precise stratigraphical allocations are given to some other localities of Central Europe. 

Until now Aplodontidae of the European Oligocene have been documented by four species only. The phylogenetic relations remained obscure. as the distribution of only one species has been known in some detail. New material made it possible to define the stratigraphic range of two of the already existing species (Plesispermophilus angustidens, Sciurodon cadurcense) and to follow their development during the Oligocene beginning with the event of the « Grande Coupure ››. Sciurodon remained nearly without change until the end of the Middle Oligocene. Plesispermophilus angustidens split into two distinct phyletic lines, one of which (P. macrodon n. sp.) reaching considerable size, is represented till the beginning of the Upper Oligocene (Pech de Fraysse, Gaimersheim). The other line leads to Plesispermophilus ernii (basal Upper Oligocene of Burgmagerbein 1. terminal Upper Oligocene of Coderet). Besides the already known forms a new small-sized species (P. atavus n. sp.) is described, which by its primitive features closely resembles the genus Plesispermophilus. Two other small-sized species already known from the Upper Oligocene (? P. argoviensis) and Lower Miocene (? P. descedens) seem to be closely related to the new species. It cannot be decided whether they are descendents of this line or have developed independently, because of their poor fossil record.
Comparison of the evolutionary modalities in the different phylogenetic lines reveals general trends. the most striking of which is the complication of the tooth pattern by the development of additional crests. In the lower molars the cusps diminuate in size and are more and more transformed into ridges. ln addition new connection between the crests appear. in the upper molars, the « selenodont » shape of the teeth becomes more and more dominant, and in the two main evolutionary lines of Plesispermophilus the metaconulus becomes duplicated. A further evolutionary trend is the size increase of the premolars compared to the molars, which is even more pronounced in the Miocene Aplodontidae.
Phylogenetic relations between the primitive Plesispermophilus and certain « prociurines ›› of Northern America as well as between Plesispermophilus (P. angustidens) and more progressive forms of the Upper Oligocene (P. ernii, P. macrodon n. sp.) can be documented. In this light, the taxonomic distinction between Prosciurinae (bunodont) and Allomyinae (selenodont) sensu Rensberger 1976 can be shown to be artificial, because it separates forms from each other, which are evidently closely related. Consequently the separation into two subfamilies has been abolished. 

The present study describes in detail the anterior limb osteology of a molossid chiropteran of the genus Tadarida, from Céreste, a Stampian locality in the Apt-Forcalquier Oligocene basin already known for its fishes, plants and insects.

A comparision with the Miocene forms T. srehlini from Saint-Géraud localities and T. sp. from Württemberg, also with the recent forms T. teniotis and Eumops perotis, does not show any clear morphological differences between the Tertiary and Recent Tadarida, indicating a rather noticeable anatomical stability, not exceptionnal indeed among Chiropterans. The Céreste fossil exhibits however slightly primitive wing proportions if compared to the Saint Gérand Aquitanian species.

Several remarks deal with the peculiar relationships between the ecology of the molossids and their kind of fossilisation, frequently associated with sedimentary facies of the lacustrine type.
  

The mammalian fauna of the Tagai locality of Olhon Island in Lake Baikal contains six artiodactyls:
Amphitragulus boulangeri, Lagomeryx parvulus, Stephanocemas sp., Palaeomeryx kaupi, Orygotherium aff. escheri, and Brachyodus intermedius. They are reported from Eastern Siberia for the first time. A description of teeth and limb-bones are given. An analysis of the systematic composition of the Tagai fauna allows it to be correlated with European faunas of the second part of the Early Miocene (Orleanian, MN 3-4) and permit suggestion of the existance of a united European-Siberian zoogeographical sub-area in the middle latitudes of Eurasia at that time. 

The Causses" is a near 3400 km2 large plateau located in the south of France. Here the first dinosaur footprints where found in 1935. After this, this area has yielded an ever-increasing number of ichnites now in excess of 500 specimens. These latter, 15 to 50 cm long, tridactyl or tetradactyl footprints of generally biped animals, were discovered at the surface of Hettangian to lower Sinemurian dolomite layers within 4 distinct stratigraphic units. The 35 sites bearing ichnites are located on the plateau margin. For the first time, morphologic characters studied through descriptive statistic methods with the usual parameters and classical Student and Snédecor tests, allowed us, to divide the whole set of biped traces into 6 ichnospecies. Their definitions are further constrained by multivariate statistical results using Principal Component Analysis (PCA), Factor Analysis of correspondances (FAC) and Discriminant Analysis (DA). All have confirmed the morphologic observations. So that now, the following taxa are identified : Grallator variabilis, G. lescurei, G. sauclierensis, G. minusculus, Eubrontes giganteus, Dilophosauripus williamsi, cf. Moraesichnium, Orníthopus fabrei nov ichnosp. The more immediately visible differences relate to the interdigital II-IV divarication and the digit length ratio. To this panel, we must add Batrachopus deweyi and shapes suggesting Trisauropodichnus and/or Anomoepus. Among all ichnite associations described in the lower Liasic, the New England assemblage presents the most affinities with ours. It shows the ichnotaxa Grallator, Dilophosauripus, Eubrontes, Batrachopus without forgetting Ornithopus fabrei nov. ichnosp. which is close to Ornithopus gallinaceus from the Massachusetts and Connecticut basins. On comparing the present early Jurassic ichnofauna of the Causses with the ones of the Middle and Upper Triassic formations of the eastem border of the Massif Central (France), it appears that tridactyl footprints become more and more numerous and large from Triassic to Early Jurassic. In the Causses, these latest are prevalent but in Quercy (France), Poland, Italy, USA, they are also associated with Omithopoda, Thyreophora and Sauropoda ichnites. Footprint areas considered here were generaly under an arid climate. Animals that passed by were heavy and bulky possible Megalosaur trackmakers, and lighter and slender Coelophysids or Ceratosaurs. For all, these areas were pathways as the orientations of the trackways seem point out. The directions followed by these reptiles were without any important variation during the Hettango-Sinemurian stages. These areas were also used from time to time by Crocodilomorpha and may be tetradactyl (I-IV) bipedal avian Theropods. However, the number of such trackways in sites, sometimes substantial, should not lead us to overestimate the trackmakers populations. These last were probably relatively moderately abondant in this inter-supratidal swamp environment. In the Causses, ichnites are connected with former algo-laminated deposits (Algal mats) which were rapidly hardened by means of calcitisation of cyanobacteria. The result has been a moderate depth of footprints; autopodia disturbing only a few cm of the carbonate substrate. Other fossils have been discovered : invertebrates with thin bivalve and gastropod shells, crustaceans tests and plants. These latter suggest the existence of paleomangroves like environments but also continental vegetation periodically overruning the swamp environment during regression/transgression cycles. At these times, wooded parts of it, could become protecting, feeding, resting and nesting places.

  

New dental and skeletal material referable to Palaeochiropteryx tupaiodon from the Middle Eocene locality of
Messel (G.F.R.) is studied, which provides additions to the previously gained knowledge of this european genus. Dental specimens from Geiseltal (G.D.R.), also of Middle Eocene age, allow us to analyze Cecilionycteria prisca. Some of these are separated to establish a new genus, Matthesia, and two new species, M. germanica and M. ? insolita. 

Cranial anatomy of a late Miocene rhizomyid, Brafhyrhizomys cf. B. pilgrimi, provides new evidence on the origin of the dorsal, round infraorbital foramen of living rhizomyines. Primitive rhizomyids retain a myomorphous keyhole foramen with a long ventral slit that retracts upward during the evolutionary history of the Rhizomyidae. The primitive condition of the elongated ventral slit is represented by Kanisamys sivalensis. Among later burrowers the foramen shows progressive dorsal migration, the ventral slit terminating midway up the snout in B.tertracharax and B. choristos ; well above the midline of the snout in Brachyrhizamys cf. B. pilgrimi. Apparently this transformation began earlier among Rhizomyinae than among Tachyoryctinae and continued to a more derived stage in rhizomyines. ln living Rhizomyx the ventral slit is absent and only a high round hole remains at the anterior end of the zygomatic arch. 

A fossil species of the extant genus Leuconoe, L. lavocati n. sp. from Le Garouillas Oligocene locality, Quercy phosphorites, SW-France, is established in nomenclatural standards. 

The object of this work is the study of the artiodactyls of the karstic locality of Lissieu near Lyon. Among the sixteen species counted, five (Hyperdichobune nobilis, Hyperdichobune sp., Haplabunodon mulleri, Tapirulus depereti and Pseudamphimeryx schlosseri) are cited for the first time in this fauna. The description of part of the anterior dentition of «Dichobune» langi permits the referal of this species to the genus Hyperdichobune. The new selenodont species Dichodon lugdunensis  described in this work belongs, with its ancestor Dichodon simplex, to a lineage distinct from that of Dichodan cartieri and Dichodon ruetimeyeri. Finally, the taxonomic reality of the Lissieu species, Lophiobunodon rhodanicum Deperet, is justified. All the other species have been compared with those of Egerkingen. Even though with most of them no difference in age can be shown between the two deposits, some however indicate clearly by their degree of evolution that Lissieu is slightly more recent than Egerkingen. ln the history of Eocene mammals, the Lissieu fauna, because of its homogeneity, can be considered as a sure reference fauna of the Auversian period. 

This paper is an English concentrate of various Russian publications by the senior author presenting the mammaIian taxa from the Cretaceous (Albian through Santonian) of the region termed Middle Asia by Soviet geographers. The diagnoses are the unmodified, literal translation of the original version, but are followed with short complementary remarks; most of the species are illuslrated anew with SEM photographs, others are by normal photography. The fossiliferous formations are cited and arguments for their dating are given. Finally, the main vertebrate groups accompanying mammaIs are listed and the environment and climate at the time of deposition are suggested. In conclusion, an hypothesis on the origin and high diversity of tribosphenic mammals on the Cretaceous coastal plains of southwest Asia is proposed. In appendix the taxon Khuduklestes bohlini novo gen. novo sp. is formally defined. 

Carolocoutoia ferigoloi gen. et sp. nov. is the largest of protodidelphid marsupials, known from Middle Paleocene levels at Itaboraí Formation, southeastern Brazil. It differs from other members of this family in having molars with low cusps which are basally inflated, rounded crests without cutting edges, and a thick enamel layer which is wrinkled, specially at the labial half. A comparative analysis among representatives of this family led us to recognize only three genera undoubtely assignable to it:  Protodidelphis PAULA COUTO, 1952, Robertbutleria MARSHALL, 1987, and Carolocoutoia gen. nov. Protodidelphids lack the basic derived features diagnostic of Polydolopimorphian marsupials, while most of its derived features agree with its belonging to the Didelphimorphia. Protodidelphids comprise a specialized clade of opossum-like marsupials adapted to frugivorous or frugivore-omnivorous feeding habits. They differ from other didelphimorphians in having very large, spire-like entoconids, reduced and antero-posteriorly compressed paraconids, absence of stylar cusp C and of para- and metaconules, large stylar cusps B and D which are proximate to each other, short postmetacristae, eccentric protocones, and molars that increase rapidly in size from M/ml to M/m3.
  

The late Pliocene site (MN 16) of Balaruc II (Hérault, France) has provided remains of the following amphibians and reptiles: Chelotriton pliocenicus nov. sp. and Triturus marmoratus (Salamandridae), cf. Rana (Ranidae), cf. Blanus (Amphisbaenidae), cf. Agama (Agamidae),  Gekkonidae indet., Lacerta s.l. (Lacertidae), "Ophisaurus" sp. (Anguidae), Michauxophis occitanus (Aniliidae), Erycinae indet. (Boidae), Elaphe cf. E. longissima and Malpolon sp. (Colubridae), cf. Naja (Elapidae) and Vipera sp. (Viperidae). The salamandrid Chelotriton pliocenicus and the aniliid Michauxophis occitanus constitute, up to now, the only records of these groups in the European Pliocene. The fauna is indicative of a warm, dry
subtropical climate. 

No abstract available 

As a result of the enrichment of eotheriodont material by one of us (P.K.T.), these specimens (essentially Biarmosuchur and Eotitanosuchur) are reexamined and refigured. A reevaluation of their particularities supports the distinction of two families, for which new diagnoses are proposed. This leads us to discuss the affinities of these families, with respect to the sphenacodonts on one hand, and to the South African primitive theriodonts on the other (gorgonopsids and ictidorhinids). This study contains inherent paleogeographic consequences which are considered in conclusion.